2021 In Paleoentomology

This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

List of years in paleoentomology

Clade Amphiesmenoptera

Lepidoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Acanthocorona

Gen. et 7 sp. nov

Valid

Mey, Léger & Lien

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Lophocoronidae moth.
The type species is A. skalskii
The genus also includes A. bowangi, A. kuranishii, A. muelleri, A. sattleri, A. spinifera, & A. wichardi.

2021 In Paleoentomology 
Acanthocorona skalskii

Agathiphagama

Gen. et sp. nov

Valid

Mey, Léger & Lien

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

An agathiphagid aglossatan moth.
The type species is A. perdita.

2021 In Paleoentomology 
Agathiphagama perdita

Phyllocnistis cretacea

Sp. nov

Valid

Fischer

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Phyllocnistis

Phyllonorycter inopinata

Sp. nov

Valid

Fischer

Lutetian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A species of Phyllonorycter

Lepidopteran research

  • The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).
  • Fossil leaf mine produced by a lyonetiid-like leaf-mining moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous (Campanian) Kaiparowits Formation (Utah, United States) by Maccracken et al. (2021), who name a new ichnotaxon Leucopteropsa spiralae.

Trichopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Archaeopolycentra achupomotet

Sp. nov

Valid

Melnitsky & Ivanov

Late Cretaceous (Santonian)

Kheta Formation
(Taymyr amber)

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Krasnoyarsk Krai)

A member of the family Polycentropodidae.

Archaeopolycentra turvalsia

Sp. nov

Valid

Melnitsky & Ivanov

Late Cretaceous (Santonian)

Kheta Formation
(Taymyr amber)

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Krasnoyarsk Krai)

A member of the family Polycentropodidae.

Cernotina fossilinova

Sp. nov

Valid

Wichard & Neumann

Probably early Miocene

Dominican amber

2021 In Paleoentomology  Dominican Republic

A caddisfly, a species of Cernotina.

Electrotrichia rasnitsyni

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Hydroptilidae.

Folindusia aenigmatosa

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Folindusia expansa

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Folindusia inconspicua

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Folindusia inventa

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Folindusia robusta

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Folindusia tortilis

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Holocentropus tutkaktut

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A species of Holocentropus.

Palaeopsilotreta kachini

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Odontoceridae.

Palaeopsilotreta succini

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Odontoceridae.

Plectrocnemia kirmikhia

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A species of Plectrocnemia.

Psilotreta fossilis

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Psilotreta.

Purbimodus khramovi

Sp. nov

Valid

Sukatsheva & Aristov

Early Cretaceous

2021 In Paleoentomology  Russia

A member of the family Vitimotauliidae.

Terrindusia nova

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

2021 In Paleoentomology  Russia

A larval case of a caddisfly.

Wormaldia denticulata

Sp. nov

In press

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A caddisfly, a species of Wormaldia.

Wormaldia diplobifurca

Sp. nov

In press

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A caddisfly, a species of Wormaldia.

Clade Antliophora

Dipterans

Name Novelty Status Authors Age Type locality Country Notes Images

Acartophthalmites crassipes

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Acartophthalmites luridus

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Acartophthalmites rugosus

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Aleksiana

Gen. et sp. et comb. nov

Valid

Krzemiński et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the crane fly family Limoniidae. The type species is A. rasnitsyni; genus also includes "Antocha" lapra Podenas & Poinar (2009).

Allarete patriciae

Sp. nov

Valid

Nel

Early Eocene

Oise amber

2021 In Paleoentomology  France

A member of the family Cecidomyiidae.

Allodia paleoafricana

Sp. nov

Valid

Bouju et al.

Early Miocene

Ethiopian amber

2021 In Paleoentomology  Ethiopia

A species of Allodia.

Apocephalus chiapanecus

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Mexican amber

2021 In Paleoentomology  Mexico

A species of Apocephalus.

Apocephalus dominicanus

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Dominican amber

2021 In Paleoentomology  Dominican Republic

A species of Apocephalus.

Apocephalus miocenus

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

A species of Apocephalus.

Araripogon pulchrus

Sp. nov

In press

Lamas, Sampronha & Ribeiro

Early Cretaceous (Aptian)

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Asilidae.

Bashkonia

Gen. et sp. nov

Valid

Lukashevich

Middle Triassic

Röt Formation

2021 In Paleoentomology  Germany

A member of the family Nadipteridae. Genus includes new species B. franconica.

Bibio castaneipennis

Sp. nov

Valid

Skartveit & Wedmann

Oligocene

2021 In Paleoentomology  Germany

A species of Bibio.

Bibio succineus

Sp. nov

Valid

Skartveit

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Bibio.

Burmahesperinus

Gen. et 3 sp. nov

Valid

Ševčík et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Bibionidae. The type species is B. antennatus; genus also includes B. conicus and B. pedicellatus.

Burmasymmerus

Gen. et 2 sp. nov

Valid

Ševčík, Hippa & Burdíková

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Ditomyiidae. The type species is B. korneliae; genus also includes B. wieslawi.

Burmatricha

Gen. et sp. nov

Valid

Ševčík, Hippa & Burdíková

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Sciaroidea of uncertain phylogenetic placement. The type species is B. mesozoica.

Cascomixticus

Gen. et sp. nov

Valid

Poinar & Vega

Cretaceous (Albian/Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Apsilocephalidae. The type species is C. tubuliferous.

Clusiomites

Fam. et gen. et comb. et sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is "Acartophthalmites" clusioides Roháček (2016); genus also includes new species C. ornatus.

Crenoptychoptera gaoi

Sp. nov

Valid

Liu & Huang

Middle–Late Jurassic

Jiyuan Basin

2021 In Paleoentomology  China

A member of the family Ptychopteridae.

Cretobibio rasnitsyni

Sp. nov

Valid

Lukashevich, Amorim & Ribeiro

Early Cretaceous (Aptian)

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Bibionidae.

Cretolimonia dayana

Sp. nov

Kopeć in Kopeć et al.

Jurassic/Cretaceous boundary

Glushkovo Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretolimonia lukashevichae

Sp. nov

Kopeć & Krzemiński in Kopeć et al.

Jurassic/Cretaceous boundary

Dain Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretolimonia mikolajczyki

Sp. nov

Kopeć, Krzemiński, Soszyńska-Maj in Kopeć et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the subfamily Architipulinae.

Cretolimonia pseudojurassica

Sp. nov

Krzemiński in Kopeć et al.

Jurassic/Cretaceous boundary

Dain Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretothereva

Gen. et sp. nov

In press

Carmo, Lamas & Ribeiro

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Therevidae. Genus includes new species C. antiqua.

Cretpenthetria

Gen. et sp. nov

Valid

Li, Zhang & Xiao

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Bibionidae. Genus includes new species C. burmensis.

Culex ekaterinae

Sp. nov

Valid

Giłka, Harbach & Perkovsky

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A mosquito, a species of Culex.

Culicoides paleopestis

Sp. nov

In press

Peñalver et al.

Eocene

Anglesea amber

2021 In Paleoentomology  Australia

A species of Culicoides.

Cyttaromyia freiwaldi

Sp. nov

Valid

Kania-Kłosok et al.

Eocene

Green River Formation

2021 In Paleoentomology  United States

A member of the family Cylindrotomidae.

Cyttaromyia gelhausi

Sp. nov

Valid

Kania-Kłosok et al.

Eocene

Green River Formation

2021 In Paleoentomology  United States

A member of the family Cylindrotomidae.

Decessia

Gen. et sp. nov

Valid

Krzemiński et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the crane fly family Limoniidae. The type species is D. podenasi.

Elephantomyia christelae

Sp. nov

Kania-Kłosok & Krzemiński

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Elephantomyia.

Elephantomyia prima

Sp. nov

Kania-Kłosok & Krzemiński

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Elephantomyia.

Forcipomyia rasnitsyni

Sp. nov

Valid

Szadziewski, Sontag & Pankowski

Miocene

Ethiopian amber

2021 In Paleoentomology  Ethiopia

A species of Forcipomyia.

Helius hispanicus

Sp. nov

Kania-Kłosok, Krzemiński & Arillo

Early Cretaceous (Albian)

Utrillas Group
(Álava amber)

2021 In Paleoentomology  Spain

A species of Helius.

Helius turolensis

Sp. nov

Kania-Kłosok, Krzemiński & Arillo

Early Cretaceous (Albian)

Utrillas Group
(San Just amber)

2021 In Paleoentomology  Spain

A species of Helius.

Hirmoneura messelense

Sp. nov

Valid

Wedmann et al.

Eocene (Lutetian)

Messel Formation

2021 In Paleoentomology  Germany

A member of the family Nemestrinidae.

Lebambromyia sacculifera

Sp. nov

Valid

Badano, Zhang, Yeates & Cerretti in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Phoroidea.

Leptotarsus andradei

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A species of Leptotarsus.

Leptotarsus gelhausi

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A species of Leptotarsus.

Leptotarsus ivoneae

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A species of Leptotarsus.

Leptotarsus lemeae

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A species of Leptotarsus.

Mesoacentron

Gen. et sp. nov

Valid

Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.

Late Cretaceous (Santonian)

Kheta Formation
(Taimyr amber)

2021 In Paleoentomology  Russia

A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is M. kaluginae.

Meunierohelea anglesensis

Sp. nov

In press

Peñalver et al.

Eocene

Anglesea amber

2021 In Paleoentomology  Australia

A member of the family Ceratopogonidae.

Minyohelea nexuosa

Sp. nov

In press

Pielowska-Ceranowska, Azar & Szwedo

Early Cretaceous (Barremian)

Lebanese amber

2021 In Paleoentomology  Lebanon

A member of the family Ceratopogonidae.

Myriophora asiatica

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Zhangpu amber

2021 In Paleoentomology  China

A species of Myriophora.

Neoarisemus groehni

Sp. nov

Valid

Wagner

Eocene

2021 In Paleoentomology  Ukraine

A member of the family Psychodidae.

Orimarguloides

Gen. et sp. nov

Valid

Men & Podenas

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the crane fly family Limoniidae. Genus includes new species O. simplex.

Palaeocentron

Gen. et sp. nov

Valid

Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.

Cretaceous (probably Albian–Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is P. krzeminskii.

Palaeoglaesum pilosus

Sp. nov

Valid

Skibińska, Krzemiński & Zhang in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Palaeoglaesum stebneri

Sp. nov

Valid

Skibińska & Krzemiński in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Palaeoglaesum teres

Sp. nov

Valid

Skibińska & Albrycht in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Paleoplatyura agnieszkae

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Paleoplatyura.

Paleoplatyura magnifica

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Paleoplatyura.

Paleoplatyura miae

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Paleoplatyura.

Paleotelmatoscopus

Gen. et comb. et sp. nov

Valid

Curler & Skibińska

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Psychodidae. The type species is "Pericoma" formosa Meunier (1905); genus also includes new species P. madrizi.

Polymera alexanderi

Sp. nov

Valid

Krzemiński et al.

Miocene

Dominican amber

2021 In Paleoentomology  Dominican Republic

A species of Polymera.

Protopenthetria

Gen. et sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Bibionidae. Genus includes new species P. skartveiti.

Rhabdomastix asiatica

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Rhabdomastix.

Rhabdomastix cretacica

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Rhabdomastix.

Rhabdomastix krzeminskae

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Rhabdomastix.

Rhabdomastix myanmae

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A species of Rhabdomastix.

Rhipidoxylomyia rasnitsyni

Sp. nov

Valid

Nel

Early Eocene

Oise amber

2021 In Paleoentomology  France

A member of the family Cecidomyiidae.

Stempellinella gilkai

Sp. nov

Valid

Zakrzewska & Jankowska

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Stempellinella.

Trichoneura xavieri

Sp. nov

Valid

Kania-Kłosok et al.

Early Cretaceous (Albian)

Utrillas Group

2021 In Paleoentomology  Spain

A species of Trichoneura.

Zhangsolva burmensis

Sp. nov

In press

Zhang & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Zhangsolvidae.

Dipteran research

  • Baranov et al. (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.

Mecopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Conicholcorpa longa

Sp. nov

Zhang, Shih & Ren in Zhang et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A scorpionfly belonging to the family Holcorpidae.

Glanomerope

Gen. et sp. nov

Valid

Poschmann & Nel

Permian

Meisenheim Formation

2021 In Paleoentomology  Germany

A scorpionfly belonging to the family Protomeropidae. Genus includes new species G. virgoferroa.

Longiphlebia incompleta

Sp. nov

In press

Lian, Cai & Huang

Late Jurassic

Tiaojishan Formation

2021 In Paleoentomology  China

A scorpionfly belonging to the family Orthophlebiidae.

Mesopsyche jinsuoguanensis

Sp. nov

Valid

Lian, Cai & Huang

Middle Triassic

Tongchuan Formation

2021 In Paleoentomology  China

A member of the family Mesopsychidae.

Mesopsyche liaoi

Sp. nov

Valid

Lian, Cai & Huang

Late Triassic

Huangshanjie Formation

2021 In Paleoentomology  China

A member of the family Mesopsychidae.

Clade Archaeorthoptera

†Caloneurodea

Name Novelty Status Authors Age Type locality Country Notes Images

Aviobiella

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Archaeorthoptera belonging to the order Caloneurodea. Genus includes new species A. garroustei.

Oudardgramma

Gen. et sp. nov

Nel & Roques

Carboniferous (Bashkirian)

2021 In Paleoentomology  France

The type species is O. bruayensis.

Paraspiculum

Gen. et sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

2021 In Paleoentomology  Germany

A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species P. brauckmannii.

†Cnemidolestodea

Name Novelty Status Authors Age Type locality Country Notes Images

Avionxixia

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is A. gui.

Ctenoptilus frequens

Sp. nov

Valid

Chen et al.

Carboniferous (Bashkirian to Moscovian)

Yanghugou Formation

2021 In Paleoentomology  China

A member of Archaeorthoptera belonging to the family Ctenoptilidae.

Palatinarkema

Gen. et sp. nov

Valid

Nel & Poschmann

Early Permian (probably Asselian–?Sakmarian)

Meisenheim Formation

2021 In Paleoentomology  Germany

A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is P. prokopi.

Piesbergopterum

Gen. et 2 sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

2021 In Paleoentomology  France
2021 In Paleoentomology  Germany

A member of Archaeorthoptera belonging to the group Cnemidolestodea. Genus includes new species P. punctatum Dvořák et al. and P. avionensis Nel & Roques.

Orthopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Amberotridactylus

Gen. et sp. nov

In press

Du, Xu & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tridactylidae. Genus includes new species A. cheni.

Archaboilus polyneurus

Sp. nov

Valid

Gu, Yue & Ren in Gu et al.

Middle Jurassic (BathonianCallovian)

Jiulongshan Formation

2021 In Paleoentomology  China

An ensiferan belonging to the group Hagloidea and the family Haglidae.

Caelielca

Gen. et sp. nov

In press

Uchida

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Elcanidae. Genus includes new species C. spinocrus.

Cascogryllus setosus

Sp. nov

In press

Du, Xu & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tridactylidae.

Cratoelcana rasnitsyni

Sp. nov

In press

Nel & Jouault

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Elcanidae.

Hukawnelca

Gen. et sp. nov

In press

Uchida

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Elcanidae. Genus includes new species H. gracile.

Panorpidium spica

Sp. nov

In press

Kim et al.

Early Cretaceous (Albian)

Jinju Formation

2021 In Paleoentomology  South Korea

An orthopteran belonging to the family Elcanidae.

Permohagla

Gen. et sp. nov

Valid

Gorochov

Late Permian

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kemerovo Oblast)

A member of Hagloidea. Genus includes new species P. tomica.

Phyllotridactylus

Gen. et sp. nov

In press

Xu et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tridactylidae. Genus includes new species P. wangi.

Pseudoacrida sennlaubi

Sp. nov

In press

Nel & Jouault

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Locustopsidae.

Orthopteran research

  • Revision of the fossil record of the family Trigonidiidae is published by Desutter-Grandcolas et al. (2021).


†Titanoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Theiatitan

Fam. et gen. et sp. nov

Valid

Schubnel, Roques & Nel in Schubnel et al.

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Titanoptera belonging to the new family Theiatitanidae. The type species is T. azari.

Other archaeorthopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Archaeogerarus

Fam. et gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Archaeorthoptera belonging to the new family Archaeogeraridae. The type species is A. schubneli.

Avionugonioneura

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Archaeorthoptera possibly belonging to the family Nugonioneuridae. Genus includes new species A. jouaulti.

Contracladus

Gen. et sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

2021 In Paleoentomology  Germany

A member of Archaeorthoptera of uncertain phylogenetic placement. Genus includes new species C. impar.

Duquesnia

Gen. et sp. nov

Valid

Nel & Roques in Nel, Roques & Duquesne

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A member of Archaeorthoptera. Genus includes new species D. gallica.

Penanoptera

Gen. et sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

2021 In Paleoentomology  Germany

A member of Archaeorthoptera. Genus includes new species P. manseri.

Clade Coleopterida

Coleoptera

Adephaga

Name Novelty Status Authors Age Type locality Country Notes Images

Antephilorhizus

Gen. et sp. nov

Valid

Kirichenko-Babko, Perkovsky & Vasilenko

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A ground beetle belonging to the tribe Lebiini. Genus includes new species A. pripiatiensis.

Balticeler

Gen. et sp. nov

Valid

Schmidt & Maddison in Schmidt, Scholz & Maddison

Eocene

Baltic amber

Europe (Baltic Sea region)

A ground beetle belonging to the subfamily Trechinae. The type species is B. kerneggeri.

Calosoma kimi

Sp. nov

Valid

Lee & Nam

Early Miocene

Geumgwangdong Formation

2021 In Paleoentomology  South Korea

A species of Calosoma.

Archostemata

Name Novelty Status Authors Age Type locality Country Notes Images

Jurodes shef

Sp. nov

Valid

Yan, Strelnikova & Cai

Early Cretaceous (Berriasian-Barremian)

Gusinoe Ozero Group

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Buryatia)

A Jurodid beetle.

Kirejtomma

Gen. et comb. nov

Valid

Li & Cai

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An Ommatid beetle
The type species is "Clessidromma" zengi (2020).

2021 In Paleoentomology 
Kirejtomma zengi

Limnomma

Gen. et sp. nov

Valid

Li & Cai in Li et al.

Middle Jurassic

Haifanggou Formation

2021 In Paleoentomology  China

An Ommatid beetle.
The type species is L. daohugouense.

2021 In Paleoentomology 
Limnomma daohugouense

Omma axsmithi

Sp. nov

In press

Jarzembowski, Zheng & Zhao

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An Omma species Ommatid beetle.

Omma forte

Sp. nov

Valid

Li & Cai
in Li, Huang & Cai

Cretaceous
(Albian to Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

An Omma species Ommatid beetle.

Paraodontomma leptocristatum

Sp. nov

Valid

Li, Yamamoto & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

An Ommatid beetle.

2021 In Paleoentomology 
Paraodontomma leptocristatum

Permocatinus

Gen. et sp. nov

Valid

Ponomarenko

Late Permian

2021 In Paleoentomology  Russia

A Coleocatiniid beetle.
The type species is P. tomiensis.

Tomiaplus

Gen. et comb. et sp. nov

Valid

Ponomarenko

Late Permian

2021 In Paleoentomology  Russia

A Triaplid beetle.
The type species is "Triaplus" sibiricus (2013),
as well as new species T. minimus.

Tunguskagyrus уani

Sp. nov

Valid

Ponomarenko

Late Permian

2021 In Paleoentomology  Russia

A Triaplid beetle.

Zygadenia alexrasnitsyni

Sp. nov

Valid

Strelnikova & Yan

Early Cretaceous

Zaza Formation

2021 In Paleoentomology  Russia

An Ommatid beetle.

Myxophaga

Name Novelty Status Authors Age Type locality Country Notes Images

Triamyxa

Fam., Gen. et sp. nov

Valid

Qvarnström et al.

Late Triassic (Carnian)

Keuper succession (Drawno Beds)

2021 In Paleoentomology  Poland

A Triamyxid beetle in the suborder Myxophaga.
The type species is T. coprolithica.

2021 In Paleoentomology 
Triamyxa coprolithica

Polyphaga

Bostrichiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Archaenosodendron

Gen. comb. et 3 sp. nov

Valid

Li & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A nosodendrid beetle.
The type species is "Nosodendron" cretaceum;
genus also includes new species A. angulare, A. explanatum and A. remotidens.

Cretomegatoma

Gen. et comb. nov

Valid

Háva

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A carpet beetle.
The type species is "Megatoma" atypica (2017).

Mesonosa

Gen. et sp. nov

Valid

Tihelka et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A wounded-tree beetle.
The type species is M. scandens.

Petalium bruteno

Sp. nov

Valid

Alekseev & Bukejs

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Petalium species death watch beetle.

Petalium widewuto

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Petalium species death watch beetle.

Trogoderma ainu

Sp. nov

Valid

Perkovsky, Háva & Zaitsev

Middle Eocene

Sakhalin amber

2021 In Paleoentomology  Russia

A Trogoderma species carpet beetle.

Tuberphradonoma

Gen. et sp. nov

Valid

Háva

Late Cretaceous
Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A carpet beetle.
The type species is T. burmitica.

Xyletinus lobanovi

Sp. nov

Valid

Bukejs, Alekseev & Háva

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Xyletinusspecies death watch beetle.

Cucujiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Aberrokorynetes oceanojubilaei

Sp. nov

In press

Kolibáč et al.

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Korynetinae checkered beetle.

Aliattalus

Gen. et sp. nov

Valid

Tshernyshev

Late Eocene

Baltic amber

2021 In Paleoentomology  Europe

A Malachiinae malachite beetle.
The type species is A. intercalaris.

Archaeosciaphilus gyrommatus

Sp. nov

In press

Legalov & Poinar

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Polydrusini Entiminae broad nosed weevil.

Arkagalarhinus

Gen. et sp. nov

In press

Legalov

Late Cretaceous
(Santonian-Campanian)

Arkagala Formation

2021 In Paleoentomology  Russia

A Chilecarinae ithycerid weevil.
The type species is A. zherikhini.

Asiomira dubrovinae

Sp. nov

Valid

Nabozhenko & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

An Alleculinae darkling beetle.

Aulacoscelis santanensis

Sp. nov

In press

Legalov

Early Cretaceous
(Aptian-Albian)

Santana Group

2021 In Paleoentomology  Brazil

An orsodacnid leaf beetle.

Baltocar sontagae

Sp. nov

Valid

Bukejs & Legalov

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Sayrevilleinae tooth-nosed snout weevil.

Baltosidis

Gen. et 3 sp. nov

Valid

Szawaryn

Eocene

Baltic amber

2021 In Paleoentomology  Europe

A Microweiseini Microweiseinae lady beetle.
The type species is B. damzeni;
genus also includes B. damgaardi and B. szadziewskii.

Bronchotibia

Gen. et sp. nov

In press

Poinar & Legalov

Miocene
(Burdigalian)

Dominican amber

2021 In Paleoentomology  Dominican Republic

A Lymantini Molytinae weevil.
The type species is B. adunatus.

Cathartosilvanus perkovskyi

Sp. nov

Valid

Alekseev & Bukejs

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A silvanid beetle.

Ceutorhynchus zerovae

Sp. nov

Valid

Legalov et al.

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A weevil.

Coslonatus

Gen. et sp. nov

Valid

Li & Cai

Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Colydiinae? Zopheridae tenebrionoid beetle.
The type species is C. rasnitsyni.

Cretaidgia

Gen. et sp. nov

In press

Zhao, Liu & Yu

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A prionocerid beetle.
The type species is C. burmensis.

Diapus ethiopicus

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Ethiopian amber

2021 In Paleoentomology  Ethiopia

A Platypodinae pinhole borer ambrosia weevil.

Diapus resinae

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Zhangpu amber

2021 In Paleoentomology  China

A Platypodinae pinhole borer ambrosia weevil.

Dorytomus mikhailovi

Sp. nov

Valid

Legalov, Nazarenko & Perkovsky

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A species of Dorytomus Curculioninae weevil.

Electroxylita

Gen. et sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A false darkling beetle.
The type species is E. chronographica.

Eurocrossotus

Gen. et sp. nov

Valid

Vitali

Eocene

Baltic amber

Europe (Baltic Sea region)

A longhorn beetle belonging to the tribe Crossotini. The type species is E. alekseevi.

Glaesotropis rohdendorfi

Sp. nov

Valid

Legalov, Nazarenko & Perkovsky

Eocene
(Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A zygaenodinin anthribine anthribid fungus weevil.

Glesoconomorphus ekaterinae

Sp. nov

Valid

Telnov et al.

Late Eocene
Priabonian

Rovno amber

2021 In Paleoentomology  Ukraine

A mycterid palm beetle.

Gracilenticrus

Gen. et sp. nov

Valid

Yu, Kolibáč & Ślipiński
in Yu et al.

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A lophocaterid beetle.
The type species is G. burmiticus.

Heterhelus buzina

Sp. nov

Valid

Kupryjanowicz, Lyubarsky & Perkovsky

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A Kateretid short-winged flower beetle. Originally described as a species of Heterhelus, but Kirejtshuk, Jenkins Shaw & Smirnov (2023) transferred it to the genus Antirhelus.

Isomira lobanovi

Sp. nov

Valid

Nabozhenko & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

An Isomira species claw comb darkling beetle.

Kaszubia

Gen. et sp. nov

Valid

Legalov, Kupryjanowicz & Perkovsky

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Poland

A cossonine weevil.
The type species is K. wanati.

Kulindrobor

Gen. et 2 sp. nov

Valid

Tihelka et al.

Late Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A tenebrionoid beetle of uncertain phylogenetic placement.
Genus includes new species K. enigmaticus and K. magnus.

Lobanoviella

Gen. et sp. nov

In press

Kirejtshuk & Reid

Eocene

Baltic amber

2021 In Paleoentomology  Europe

A lobanovielline palophagine megalopodid leaf beetle.
The type species is L. andreyi.

Madelinia capillata

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A false darkling beetle.

Melanosiagon

Gen. et sp. nov

Valid

Batelka & Prokop

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A wedge-shaped beetle.
The type species is M. serraticornis.

Metaclisa ottoi

Sp. nov

Valid

Nabozhenko, McKellar & Bukejs

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

AMetaclisa species metaclisine darkling beetle.

Microscapha kugelanni

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A false darkling beetle.

Microtrogossita

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Trogossitid bark-gnawing beetle.
The type species is M. qizhihaoi.

Monolepta rappsilberi

Sp. nov

Valid

Bukejs et al.

Late Eocene
Priabonian

Bitterfeld amber

2021 In Paleoentomology  Germany

A Monolepta species skeletonizing leaf beetle.

Neolatitergum

Nom. nov

Valid

Zhang

Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A Trogossitid bark-gnawing beetle.
A replacement name for "Latitergum" Yu et al. (2014).

Oedemera girulskii

Sp. nov

Valid

Szawaryn, Sontag & Kubisz

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

An Oedemera species false blister beetle.

Pacyclaxyra

Gen. et sp. nov

Valid

Tihelka, Huang & Cai

Late Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A sooty mould beetle.
The type species is P. azari.

Palaeatalasis

Gen. et sp. nov

Valid

Legalov

Eocene
(Ypresian)

Green River Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Utah)

A Megamerini Sagrinae leaf beetle.
The type species is P. monrosi.

Palaeoanoplus

Trib., Gen., et sp. nov

Valid

Legalov

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Palaeoanoplini Curculioninae weevil.
The type species is P. horridus.

Paleopegorhinus

Gen. et sp. nov

Valid

Poinar, Brown & Legalov

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A Chilecarinae new york weevil.
The type species is P. micrommatus.

Palaeopeplus

Gen. et sp. nov

Valid

Powell & Cline

Miocene
(Burdigalina)

Dominican amber

2021 In Paleoentomology  Dominican Republic

A Cillaeinae sap beetle.
The type species is P. cascus.

Pelretes

Gen. et sp. nov

Valid

Tihelka et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A short-winged flower beetle.
The type species is P. vivificus .

Phloiotrya inmarinata

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Phloiotrya species false darkling beetle.

Protodasytes

Gen. et sp. nov

Valid

Tihelka et al.

Late Cretaceous
(Cenomanian)

Charentese amber

2021 In Paleoentomology  France

A soft-wing flower beetle.
The type species is P. cretaceus.

Protomauroania

Gen. et 2 sp. nov

Valid

Tshernyshev in Tshernyshev & Shcherbakov

Late Eocene

Baltic amber
Rovno amber

Baltic Sea region
2021 In Paleoentomology  Ukraine

A Dasytinae malachite beetle.
The type species P. chaetophorata
Genus also includes P. mikhailovi.

Protostomopsis

Gen. et sp. nov

Valid

Bukejs, Ślipiński & Alekseev
in Bukejs et al.

Eocene

Baltic amber

2021 In Paleoentomology  Poland
2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A cerylonid beetle. The type species is P. pandema.

Pseudomesauletes lobanovi

Sp. nov

Valid

Bukejs & Legalov

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Rhynchitinae tooth-nosed snout weevil.

Pulchritudo

Gen. et sp. nov

Valid

Krell & Vitali

Eocene
(Ypresian)

Green River Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Colorado)

A Sagrinae leaf beetle.
The type species is P. attenboroughi.

Rhamphophorus

Trib., gen. et sp. nov

Valid

Poinar & Brown

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Rhamphophorini Cimberidinae pine flower weevil.
The type species is R. legalovii.

Salignacicola

Gen. et comb. nov

Valid

Batelka & Prokop

Late Cretaceous
(Cenomanian)

2021 In Paleoentomology  France

A wedge shaped beetle.
The type species is "Macrosiagon" ebboi (2004).

Symphora glaesonauta

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A false darkling beetle.

Symphora pollocki

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A false darkling beetle.

Teredolaemus primigenius

Sp. nov

Valid

Alekseev et al.

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A teredid beetle.

Toxorhynchus michalskii

Sp. nov

Valid

Legalov

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Apioninae straight-snouted weevil.

Trematosphindus

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A slime mold beetle.
The type species is T. newtoni

Zaiwa

Gen. et sp. nov

Valid

Lyubarsky et al.

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A lophocaterid beetle.
The type species is Z. pankowskiorum.

Zherikhiniellus

Gen. et sp. nov

Valid

Legalov

late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Nanophyinae straight-snouted weevil.
The type species is Z. rarus.

Cucujiformia research
  • A piece of Burmese amber preserving the complete series of free-living stages (except of the last larval instar) of a Cretaceous parasitoid beetle Paleoripiphorus is described by Batelka, Engel & Prokop (2021).
Elateriformia
Name Novelty Status Authors Age Type locality Country Notes Images

Amberophytum maculatum

Sp. nov

Valid

Poinar, Vega & Legalov

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cerophytid rare click beetle.

Anoeuma

Gen. et sp. nov

Valid

Li, Kundrata & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

An elateroid click beetle of uncertain phylogenetic placement.
The type species is A. lawrencei.

Arturmiles

Gen. et sp. nov

Valid

Fanti

Eocene

Prussian Formation
(Baltic amber)

2021 In Paleoentomology  Poland

A soldier beetle belonging to the tribe Cantharini. The type species is A. pankowskiarum.

Balistica vetai

Sp. nov

Valid

Muona

Eocene

Baltic amber

2021 In Paleoentomology  Europe

A Dirhagini Melasinae false click beetle.

Baltodascillus

Gen. et sp. nov

Valid

Kundrata et al.

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A dascillid beetle.
The type species is B. serraticornis.

Burmagrilus

Gen. et sp. nov

Valid

Jiang, Song & Wang in Jiang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A metallic wood-boring beetle.
The type species is B. cretacus.

Burmomiles bilineatimaculatus

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.

Burmomiles laticollis

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle.

Burmomiles oblongoculus

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.

Captopus

Gen. et sp. nov

Valid

Li & Cai in Li, Huang & Cai

Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A throscid beetle.
The type species is C. depressiceps.

Chelonarium andabata

Sp. nov

Valid

Alekseev & Bukejs in Alekseev et al.

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A turtle beetle.

Chelonarium dingansich

Sp. nov

Valid

Alekseev & Bukejs in Alekseev et al.

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A turtle beetle.

Chelonarium dominicanum

Sp. nov

Valid

Poinar & Háva

Burdigalian

Dominican amber

2021 In Paleoentomology  Dominican Republic

A turtle beetle.

Coomanius balticus

Sp. nov

Valid

Muona

Eocene

Baltic amber

2021 In Paleoentomology  Europe

A Nematodini Macraulacinae false click beetle.

Cretocantharis

Gen. et sp. nov

Valid

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle.
The type species is C. veda.

Cretopachyderes

Gen. et sp. nov

Valid

Kundrata et al.

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

An Agrypninae click beetle.
The type species is C. burmitinus.

Cretophengodes

Fam. nov., Gen. et sp. nov

Valid

Li, Kundrata, Tihelka & Cai in Li et al.

Cretaceous (Albian or Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cretophengodid elateroid beetle.
The type species is C. azari.

Dyscharachthis frickae

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe (Baltic Sea region)

A Dyscharachthini Eucneminae false click beetle.

Electrothroscus

Gen. et sp. nov

Valid

Li & Cai

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A throscid false click beetle.
The type species is E. yanpingae.

Entomophthalmus antennalis

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe (Baltic Sea region)

A Dirhagini Melasinae false click beetle.

Erdaia bispinulosa

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Orodotini? Macraulacinae false click beetle.

Erdaia longicornis

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Orodotini? Macraulacinae false click beetle.

Erdaia longitarsis

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Orodotini? Macraulacinae false click beetle.

Euryptychoides

Gen. et sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.
The type species is E. magnus.

Euryptychus decoratus

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus excavatus

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus helmwigae

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus sieglindae

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus waltrautae

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Euryptychini Macraulacinae false click beetle.

Hukawngichthyurus maha

Sp. nov

Valid

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle.

Isorhipis balticus

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

An Isorhipis species melasine false click beetle

Lycocerus crassepunctatus

Sp. nov

Valid

Kazantsev

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A soldier beetle.

Malthodes aliger

Sp. nov

Valid

Kazantsev

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A soldier beetle

Malthodes headsi

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Poland

A soldier beetle.

Malthodes jaredi

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Poland

A soldier beetle, a species of Malthodes.

Malthodes vladiceniensis

Sp. nov

Valid

Pintilioaie, Fanti & Ionesi

Middle Miocene

Bârnova-Muntele Formation

2021 In Paleoentomology  Romania

A soldier beetle.

Microrhagus (Emyirhagus) vetai

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Microrhagusspecies melasine false click beetle

Murcybolus

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Burmolycini dexorine net-winged beetle.
The type species is M. longiantennus.

Nematodes schwertleitae

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Nematodini Macraulacinae false click beetle.

Neocharoides

Gen. et sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Nematodini? Macraulacinae false click beetle.
The type species is N. balticus.

Palaeocantharis

Gen. et sp. nov

Disputed

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A soldier beetle. The type species is P. panna.
Fanti & Müller (2022) considered the genus Palaeocantharis to a be a junior synonym of the genus Elektrokleinia, though the authors maintained P. panna as a distinct species within the latter genus.

Podistra guthriei

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Poland

A soldier beetle.

Poecilochrus balticus

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Eucnemini Eucneminae false click beetle.

Potergites brittoni

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Dendrocharini Eucneminae false click beetle.

Pseudomataeopsephus

Gen. et sp. nov

In press

Li, Huang & Cai

Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A water-penny beetle.
The type species is P. burmensis.

Pseudopactopus

Gen. et sp. nov

Valid

Li & Cai
in Li, Huang & Cai

Cretaceous
( Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A throscid false click beetle.
The type species is P. robustus.

Ptilodactyla eocenica

Sp. nov

Valid

Kundrata, Bukejs & Blank
in Kundrata et al.

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

APtilodactyla species toe winged beetle.

Rhacopus balticus

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Rhacopus species eucnemid false click beetle

Rhagomicrus balticus

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Dirhagini Melasinae false click beetle.

Rheanischia

Gen. et sp. nov

Li, Chang & Muona

Early Cretaceous
(Hauterivian to Aptian)

Yixian Formation

2021 In Paleoentomology  China

An Anischiinae false click beetle.
The type species is R. brevicornis.

Silis curleri

Sp. nov

Valid

Fanti & Pankowski

Burdigalian

Dominican amber

2021 In Paleoentomology  Dominican Republic

A species of Silis.

Silis hegnai

Sp. nov

Valid

Fanti & Pankowski

Burdigalian

Dominican amber

2021 In Paleoentomology  Dominican Republic

A species of Silis.

Xylophilus balticus

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A Xylophilus species eucnemid false click beetle.

Elateriformia research
  • A study on the anatomy and phylogenetic relationships of Archaeolus funestus is published by Li et al. (2021).
  • A study on the morphological diversity of larvae of extant and fossil members of the family Brachypsectridae is published by Haug et al. (2021).
Scarabaeiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Goriresina

Gen. et sp. nov

Valid

Matalin, Perkovsky & Vasilenko

Late Eocene
Priabonian

Rovno amber

2021 In Paleoentomology  Ukraine

A tiger beetle.
The type species is G. fungifora.

Onthophagus pilauco

Sp. nov

Valid

Tello et al.

Late Pleistocene

Pilauco Bajo

2021 In Paleoentomology  Chile

A scarab beetle.

Staphyliniformia
Name Novelty Status Authors Age Type locality Country Notes Images

Acritus sutirca

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A species of Acritus.

Bacanius gorskii

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Poland

A species of Bacanius.

Burmagluta

Gen. et sp. nov

Valid

Yin & Cai

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A Brachyglutini Pselaphinae rove beetle.
The type species is B. rougemonti.

Carinumerus

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An Onthophilinae clown beetle.
The type species is C. maddisoni.

Charhyphus serratus

Sp. nov

In press

Yamamoto & Shavrin

Eocene

Baltic amber

2021 In Paleoentomology  Europe

A Phloeocharinae rove beetle.

Cretosaja

Gen. et sp. nov

In press

Sohn & Nam

Early Cretaceous

Jinju Formation

2021 In Paleoentomology  South Korea

A carrion beetle.
The type species is C. jinjuensis.

Druantia

Gen. et sp. nov

Valid

Caterino

Cenomianian

Burmese amber

2021 In Paleoentomology  Myanmar

A Dendrophilinae clown beetle.
The type species is D. aeterna.

Euconnus nathani

Sp. nov

Valid

Jałoszyński & Perkovsky

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A Euconnus species Scydmaeninae rove beetle.

Eusphalerum bukejsi

Sp. nov

Valid

Shavrin in Shavrin & Kairišs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Eusphalerum species Omaliinae rove beetle.

Megabythinus

Gen. et sp. nov

Valid

Yin, Zhao & Cai

Cretaceous
Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A Bythinini Pselaphinae rove beetle.
The type species is M. spinitibialis.

Paroxyporus

Gen. et sp. nov

Valid

Cai

Early Cretaceous

Yixian Formation

2021 In Paleoentomology  China

A rove beetle.
The type species is P. fossilis.

Phasmister

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An Onthophilinae clown beetle.
The type species is P. cristatus.

Ponohydrochus

Gen. et sp. nov

Valid

Prokin & Strelnikova

Early Cretaceous
(Hauterivian)

2021 In Paleoentomology  Russia

A water scavenger beetle.
The type species is P. buryaticus.

Proptomaphaginus alleni

Sp. nov

Valid

Perreau & Haelewaters
in Perreau, Haelewaters & Tafforeau

Miocene
Burdigalian

Dominican amber

2021 In Paleoentomology  Dominican Republic

A Ptomaphagini round fungus beetle.

Trichophya minor

Sp. nov

Valid

Yamamoto & Newton

Cretaceous
(Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A Trichophya species Trichophyinae rove beetle.

Yethiha

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A Dendrophilinae clown beetle.
The type species is Y. peregrina.

Coleopteran research

  • A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the Permian–Triassic extinction event on beetle diversity, is published by Zhao et al. (2021).
  • Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus Notocupes, transfer N. ? multituberatus to the genus Lasiosyne, and consider "N." undatabdominus to be likely a member of the family Artematopodidae.
  • The oldest solid wood-borer larvae reported to date (belonging to the families Buprestidae and possibly Cerambycidae) are described from the Cretaceous amber from Myanmar by Haug et al. (2021).
  • Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from pack rat middens from California (United States) and Baja California (Mexico) dating up to ~ 34,355 years before present by Smith et al. (2021).

Clade Dictyoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Afrophaga

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A cockroach belonging to the family Corydiidae. Genus includes new species A. extincta.

Alienopterix mlynskyi

Sp. nov

In press

Sendi in Vršanský et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An alienopterid.

Alienopterix smidovae

Sp. nov

In press

Hinkelman in Vršanský et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An alienopterid.

Arceotermes

Fam. et gen. et sp. nov

Valid

Engel & Jiang in Jiang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A termite belonging to the new family Arceotermitidae. Genus includes new species A. hospitis.

Caligoptera

Gen. et sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Ectobiidae. Genus includes new species C. hinkelmani.

Chuanblatta

Gen. et 2 sp. nov

In press

Liang et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A cockroach belonging to the family Raphidiomimidae. Genus includes new species C. huntianlingensis and C. lata.

Cuniculoblatta

Gen. et sp. nov

In press

Hinkelman

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach. Genus includes new species C. brevialata .

Ectoovia

Gen. et sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Ectobiidae. Genus includes new species E. protecta.

? Elisama algeriaensis

Sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of the family Blattulidae.

Eminespina

Gen. et sp. nov

Valid

Chen, Zhang Shi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

An alienopterid. Genus includes new species E. burma.

Fractalia

Fam. et gen. et 2 sp. nov

Valid

Hinkelman & Vršanský in Vršanský et al.

Late Jurassic–Late Cretaceous (Kimmeridgian–Cenomanian)

Burmese amber

2021 In Paleoentomology  Germany
2021 In Paleoentomology  Kazakhstan
2021 In Paleoentomology  Myanmar

A member of Corydioidea, the type genus of the new family Fractaliidae Vršanský & Hinkelman. The type species is F. articulata Hinkelman; genus also includes new species F. aristovi Vršanský.

Fragosublatta

Gen. et sp. nov

Valid

Chen, Shih & Ren in Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Corydiidae. The type species is F. pectinata.

Glyptotermes abyssinicus

Sp. nov

Valid

Bouju, Jouault & Perrichot

Early Miocene

Ethiopian amber

2021 In Paleoentomology  Ethiopia

A termite belonging to the family Kalotermitidae.

Kinneyblatta

Gen. et sp. nov

Valid

Schneider et al.

Carboniferous (Kasimovian)

Atrasado Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  New Mexico)

A member of Blattodea belonging to the family Spiloblattinidae. The type species is K. huberi.

Kriedoblatta

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of the family Liberiblattinidae. Genus includes new species K. gondwanensis.

? Kurablattina samsonovi

Sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of the family Liberiblattinidae.

Labradormantis

Gen. et sp. nov

Valid

Demers‐Potvin et al.

Late Cretaceous (Cenomanian)

Redmond Formation

2021 In Paleoentomology  Canada

A praying mantis belonging to the family Baissomantidae. Genus includes new species L. guilbaulti.

Latiblattella basaltica

Sp. nov

In press

Vršanský, Poschmann & Vidlička

Late Oligocene

2021 In Paleoentomology  Germany

A species of Latiblattella.

Latiblattella karlgruberi

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

Laticephalana

Gen. et sp. nov

Valid

Luo et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Umenocoleidae. Genus includes new species L. liuyani.

Lukotermes

Gen. et sp. nov

Valid

Perkovsky & Nel

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A termite belonging to the family Rhinotermitidae. Genus includes new species L. milescaput.

Meiatermes cretacicus

Sp. nov

In press

Bezerra et al.

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A termite.

Meloblatta

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of the family Mesoblattinidae. Genus includes new species M. brezinica.

Milesitermes

Gen. et sp. nov

Valid

Jouault et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A termite belonging to the family Mastotermitidae. Genus includes new species M. engeli.

Miniblattina

Gen. et sp. nov

In press

Sendi

Early Cretaceous (Barremian)

Lebanese amber

2021 In Paleoentomology  Lebanon

A cockroach belonging to the family Liberiblattinidae. Genus includes new species M. libera.

Mongolblatta sendii

Sp. nov

In press

Hinkelman

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Mesoblattinidae.

Morphna cenozoica

Sp. nov

In press

Šmídová, Vidlička & Wedmann

Eocene

Messel pit

2021 In Paleoentomology  Germany

A cockroach belonging to the family Blaberidae.

Morphna cretacica

Sp. nov

In press

Šmídová, Vidlička & Wedmann

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Blaberidae.

Nadveruzenie

Gen. et sp. nov

In press

Vršanský, Hinkelman & Sendi in Vršanský et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An alienopterid. Genus includes new species N. postava.

Nodosigalea simplivena

Sp. nov

In press

Šmídová

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Corydiidae.

Ocelloblattula margarita

Sp. nov

Valid

Koubová & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Corydioidea.

Olenablatta

Gen. et sp. nov

In press

Šmídová

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Olidae. Genus includes new species O. vrsanskyi.

Otazka

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of the family Mesoblattinidae. Genus includes new species O. systematicka.

Pabuonqed magna

Sp. nov

In press

Song et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Mastotermitoidea belonging to the family Pabuonqedidae.

Paekthoblatta

Gen. et sp. nov

Valid

So et al.

Early Cretaceous

Sinuiju Formation

2021 In Paleoentomology  North Korea

A member of the family Raphidiomimidae. Genus includes new species P. coreanica.

Periplaneta (?) perialla

Sp. nov

Valid

Anisyutkin & Perkovsky

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A cockroach, possibly a species of Periplaneta.

Petropterix fukuiensis

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

2021 In Paleoentomology  Japan

A member of the family Umenocoleidae.

Praeblattella arcuata

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

2021 In Paleoentomology  Japan

A member of the family Mesoblattinidae.

Praeblattella inexpecta

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

2021 In Paleoentomology  Japan

A member of the family Mesoblattinidae.

Pseudomantina

Gen. et 2 sp. nov

Valid

Sendi & Vršanský in Vršanský et al.

Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)

Lebanese amber

2021 In Paleoentomology  Kazakhstan
2021 In Paleoentomology  Lebanon

A member of the family Liberiblattinidae. The type species is P. occisor Sendi; genus also includes new species P. nigroalba Vršanský.

Reticulitermes grimaldii

Sp. nov

Valid

Jouault et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A termite, a species of Reticulitermes.

Sajda

Gen. et sp. nov

Junior homonym

Vršanský et al.

Cretaceous (Hauterivian?)

2021 In Paleoentomology  Algeria

A member of Holocompsinae. Genus includes new species S. equatorialis. The generic name is preoccupied by Sajda Dworakowska (1981); Jiang, Xing & Li (2023) coined a replacement name Vrsanskysajda.

Spinaeblattina baekthoensis

Sp. nov

Valid

So et al.

Early Cretaceous

Sinuiju Formation

2021 In Paleoentomology  North Korea

A member of the family Mesoblattinidae.

Stavba delicata

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Liberiblattinidae.

Stavba magnoculara

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A cockroach belonging to the family Liberiblattinidae.

Tanytermitalis

Gen. et sp. nov

Valid

Engel & Cai in Jiang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A termite belonging to the family Tanytermitidae. Genus includes new species T. philetaerus.

Dictyopteran research

  • A study on the phylogenetic placement of the family Umenocoleidae, based on data from a new specimen of Umenocoleus sinuatus from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo et al. (2021).
  • Description of new alienopterid nymphs from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo et al. (2021).
  • A study on the sensory organs of Huablattula hui, and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi et al. (2021).

Clade †Glosselytrodea

Name Novelty Status Authors Age Type locality Country Notes Images

Moltenojurina

Gen. et sp. nov

Valid

Béthoux & Anderson

Late Triassic (Carnian)

Molteno Formation

2021 In Paleoentomology  South Africa

A polycytellid Glosselytrodean.
The type species is M. parva.

Polycytella rasnitsyni

Sp. nov

Valid

Béthoux & Anderson

Late Triassic (Carnian)

Molteno Formation

2021 In Paleoentomology  South Africa

A polycytellid Glosselytrodea.

Hymenopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Aethemelikertes

Gen. et sp. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is A. emunctorii.

Alivespa gracilenta

Sp. nov

In press

Wu et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Vespidae.

Allommation

Fam. nov., Gen. et 3 sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Apoidea in the new family Allommationidae. Genus includes new species A. procerum, A. elongatum and A. grande.

Amelikertotes

Gen. et comb. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber
Bitterfeld amber

2021 In Paleoentomology  Germany
2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is "Melikertes" clypeatus Engel (2001).

Ammosphecium

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Crabroninae. Genus includes new species A. diabolicum.

Anochetus miosumatrensis

Sp. nov

Valid

Ngô-Muller, Garrouste & Nel in Ngô-Muller et al.

Miocene

2021 In Paleoentomology  Indonesia

An ant, a species of Anochetus.

Archaeovespa incompleta

Sp. nov

In press

Wu et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Vespidae.

Archeonesia

Gen. et sp. nov

Valid

Tribull, Pankowski & Colombo

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A Pristocerinae Bethylid.
The type species is A. eocena.

2021 In Paleoentomology 
Archeonesia eocena

Arkadiserphus

Gen. et sp. nov

Valid

Rasnitsyn

Late Jurassic

Karabastau Formation

2021 In Paleoentomology  Kazakhstan

A member of Proctotrupoidea belonging to the family Peleserphidae. The type species is A. leleji.

Ascogaster (Syntaphus) latitibialis

Sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Cheloninae.

Aulacicosmus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Heterogynaidae. Genus includes new species A. setosus.

Austroplebeia fujianica

Sp. nov

Valid

Engel in Engel et al.

Miocene (Langhian)

Zhangpu amber

2021 In Paleoentomology  China

A stingless bee, a species of Austroplebeia.

Burmabracon

Gen. et 2 sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Braconidae belonging to the subfamily Protorhyssalinae. Genus includes new species B. gracilens and B. grossus.

Burmapyris

Gen. et sp. nov

Valid

Jouault, Perrichot & Nel

Cretaceous

2021 In Paleoentomology  Myanmar

A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species B. azevedoi.

Burmoxyela

Gen. et sp. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A sawfly belonging to the family Syspastoxyelidae. Genus includes new species B. lii.

Burmusculus angustus

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Apocrita belonging to the family Burmusculidae.

Buserphites

Gen. et 2 sp. nov

In press

Herbert & McKellar

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Serphitidae. Genus includes new species B. applanatus and B. myanmarensis.

Chainochora

Gen. et sp. nov

Valid

Chen, van Achterberg & Hong

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is C. syntoma.

Chlorepyris deploegi

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Bethylidae.

Chlorepyris engeli

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae.

Chlorepyris hopei

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae.

Chlorepyris jouaulti

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Bethylidae.

Chlorepyris mckellari

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Bethylidae.

Chlorepyris meunieri

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae.

Chlorepyris neli

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber
Rovno amber

2021 In Paleoentomology  Russia
2021 In Paleoentomology  Ukraine

A member of the family Bethylidae.

Chlorepyris perrichoti

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae.

Christophus

Gen. et 2 sp. nov

In press

Brazidec et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Diapriidae. Genus includes new species C. myanmarensis and C. ohmkuhnlei.

Cretastatus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Astatinae. Genus includes new species C. longevus.

Cretorussus

Gen. et sp. nov

In press

Jouault, Perrichot & Nel

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Orussoidea. Genus includes new species C. vilhelmseni.

Curiosivespa sennlaubi

Sp. nov

Valid

Jouault, Rasnitsyn & Nel in Jouault et al.

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Vespidae.

Diameneura

Gen. et sp. nov

In press

Santer & Álvarez-Parra in Santer et al.

Early Cretaceous (Albian)

Escucha Formation

2021 In Paleoentomology  Spain

A member of the family Spathiopterygidae. The type species is D. marveni.

Drymomyrmex rasnitsyni

Sp. nov

Valid

Radchenko

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

An ant belonging to the subfamily Formicinae.

Elasmosomites arkadyleleji

Sp. nov

Valid

Belokobylskij in Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Euphorinae.

Electrofoenops exaltatus

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Aulacidae.

Embolemus brachypterus

Sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Embolemidae.

Embolemus micropterus

Sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Embolemidae.

Eopelecinus marechali

Sp. nov

Valid

Jouault

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Pelecinidae.

Eoteredon

Gen. et sp. nov

Valid

Archibald, Aase & Nel

Early Eocene

Green River Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Wyoming)

A horntail. Genus includes new species E. lacoi.

Ephedrus rasnitsyni

Sp. nov

Valid

Davidian & Kaliuzhna in Davidian, Kaliuzhna & Perkovsky

Middle Eocene

Sakhalinian amber

2021 In Paleoentomology  Russia

A member of the family Braconidae belonging to the subfamily Aphidiinae.

Exilaulacus advenus

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Aulacidae.

Exilaulacus eximius

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Aulacidae.

Formica ribbeckei

Sp. nov

Valid

Radchenko & Perkovsky in Radchenko, Perkovsky & Vasilenko

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

An ant, a species of Formica.

Glenocephalus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Apoidea in the new family Cirrosphecidae. Genus includes new species G. mandibularis.

Gloxinius

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae; a new genus for "Epyris" bifossatus Brues (1939).

Grandixyela

Gen. et sp. nov

Valid

Zheng in Zheng et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Syspastoxyelidae. The type species is G. rasnitsyni.

Haidomelikertes

Gen. et sp. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is H. uraeus.

Haptodioctes

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Apoidea in the new family Cirrosphecidae. Genus includes new species H. apiformis.

Helosericus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Psenini. Genus includes new species H. verrucosus.

Heteroichneumon

Gen. et sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Ichneumonidae. The type species is H. rasnitsyni.

Heteropimpla pulverulenta

Sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Ichneumonidae.

Holepyris rasnitsyni

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae.

Holepyris terayamai

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Bethylidae.

Holopsenella antiqua

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Holopsenella burmitica

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Holopsenella gothica

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Hybristodryinus castaneus

Sp. nov

Valid

Olmi, Guglielmino & Chen in Olmi et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Dryinidae.

Hybristodryinus viriosus

Sp. nov

Valid

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Dryinidae.

Hybristodryinus zaifui

Sp. nov

Valid

Chen, Olmi & Perkovsky in Olmi et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Dryinidae.

Janzenella theia

Sp. nov

Valid

Bremer & Talamas in Bremer, van de Kamp & Talamas

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A species of Janzenella.

Karataus ryonsangensis

Sp. nov

In press

Won & So

Early Cretaceous (BarremianAptian)

Sinuiju Formation

2021 In Paleoentomology  North Korea

A wasp belonging to the family Ephialtitidae.

Keratodellitha

Gen. et 3 sp. nov

Valid

Jouault et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Evanioidea belonging to the family Othniodellithidae. Genus includes new species K. anubis, K. basilisci and K. kirin.

Kleistochora

Gen. et sp. nov

Valid

Chen, van Achterberg & Hong

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is K. dolichura.

Lelejus

Gen. et sp. nov

Valid

Radchenko & Proshchalykin

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

An ant belonging to the subfamily Myrmicinae. The type species is L. venustus.

Lonchodryinus groehni

Sp. nov

Valid

Olmi et al.

Late Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Dryinidae.

Mael

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

2021 In Paleoentomology  Russia

A member of the family Bethylidae belonging to the subfamily Scleroderminae; a new genus for "Calyoza" longiceps Brues (1923).

Megalyrhyssalus

Gen. et sp. nov

Valid

Belokobylskij & Jouault

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Braconidae. Genus includes new species M. clavicornis.

Megarolium

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Bembicinae. Genus includes new species M. paradoxum.

Merascylla

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Bethylidae belonging to the subfamily Pristocerinae; a new genus for "Epyris" atavellus Cockerell (1920).

Mesoserphites

Gen. et 5 sp. nov

In press

Herbert & McKellar

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Serphitidae. Genus includes new species M. annulus, M. giganteus, M. engeli, M. scutatus and M. viraneacapitis.

Meteorus arasnitsyni

Sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Meteorus.

Microtypus eocenus

Sp. nov

Valid

Belokobylskij et al.

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Braconidae belonging to the subfamily Microtypinae.

Mintara

Gen. et sp. nov

In press

Brazidec et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Diapriidae. Genus includes new species M. parva.

Odontosericus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Psenini. Genus includes new species O. burmensis.

Orthosyntexis

Gen. et 2 sp. nov

Valid

Gao, Engel, Shih, Gao in Gao et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Anaxyelidae. The type species is O. elegans; genus also includes O. thanti.

Palaeocolastes

Gen. et sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Exothecinae. The type species is P. bruesi.

Paleoaulacus

Gen. et sp. nov

Valid

Jouault & Nel

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Aulacidae. Genus includes new species P. minutus.

Paleoxyela

Gen. et sp. nov

Valid

Jouault, Aase & Nel

Eocene (Chadronian)

Florissant Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Colorado)

A member of the family Xyelidae. The type species is P. nearctica.

Peleserphus qufuensis

Sp. nov

In press

Yu et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Peleserphidae.

Pheidole pauchil

Sp. nov

Valid

Varela-Hernández & Riquelme

Miocene

Mexican amber

2021 In Paleoentomology  Mexico

An ant, a species of Pheidole.

Pinguixyela

Gen. et comb. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A sawfly belonging to the family Syspastoxyelidae; a new genus for "Syspastoxyela" pinguis.

Platyxyela tenuis

Sp. nov

Valid

Zheng et al.

Middle Jurassic (Callovian-Bathonian boundary)

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Xyelidae.

Pompilopterus ryonsangensis

Sp. nov

In press

Won & So

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

2021 In Paleoentomology  North Korea

A wasp belonging to the family Angarosphecidae.

Pristinopterus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Bembicinae. Genus includes new species P. spectabilis.

Pristopsen

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Psenini. Genus includes new species P. obscurus.

Prosyntexis sennlaubi

Sp. nov

Valid

Jouault & Nel

Early Cretaceous (Aptian)

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Sepulcidae.

Protopyris

Gen. et sp. nov

Valid

Jouault & Nel in Jouault, Perrichot & Nel

Early Cretaceous (Albian)

Hkamti amber

2021 In Paleoentomology  Myanmar

A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species P. myanmarensis.

Prototeleia

Gen. et sp. nov

Valid

Talamas et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Platygastridae. The type species is P. kleio.

Proxyelia

Gen. et sp. nov

Valid

Jouault, Aase & Nel

Eocene (Ypresian)

Green River Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Wyoming)

A member of the family Xyelidae. The type species is P. pankowskii.

Ptilocosmus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Heterogynaidae. Genus includes new species P. corniculatus.

Ramageoptera

Gen. et sp. nov

Valid

Jouault & Brazidec

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Bethylidae belonging to the subfamily Protopristocerinae. Genus includes new species R. platycephala.

Rasnichneumon

Gen. et 2 sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Ichneumonidae. The type species is R. alexandri; genus also includes R. gracilis.

Rasnitsynum

Gen. et sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Dryinidae. Genus includes new species R. burmense.

Rogichneumon

Gen. et sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Ichneumonidae. The type species is R. braconidicus.

Rovenosa alexrasnitsyni

Sp. nov

Valid

Manukyan in Manukyan & Zhindarev

Late Eocene

Prussian Formation
(Baltic amber)

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of the family Ichneumonidae.

Rovenosa khalaimi

Sp. nov

Valid

Manukyan in Manukyan & Zhindarev

Late Eocene

Prussian Formation
(Baltic amber)

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A member of the family Ichneumonidae.

Sakhalinencyrtus

Gen. et sp. nov

Valid

Simutnik in Simutnik, Perkovsky & Vasilenko

Middle Eocene

Sakhalinian amber

2021 In Paleoentomology  Russia

A member of the family Encyrtidae. The type species is S. leleji.

Sakhalinoctonus

Gen. et sp. nov

Valid

Davidian in Davidian, Manukyan & Belokobylskij

Middle Eocene

Sakhalinian amber

2021 In Paleoentomology  Russia

A member of the family Braconidae belonging to the subfamily Aphidiinae. Genus includes new species S. alexrasnitsyni.

Scleroxyela

Gen. et sp. nov

Valid

Zheng et al.

Middle Jurassic (Callovian-Bathonian boundary)

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Xyelidae. The type species is S. daohugouensis.

Spalangiopelta darlingi

Sp. nov

Valid

Moser in Moser et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Pteromalidae.

Spalangiopelta semialba

Sp. nov

Valid

Moser in Moser et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Pteromalidae.

Spheciellus

Fam. nov., Gen. et sp. nov.

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Apoidea in the new family Spheciellidae. Genus includes new species S. aenigmaticus.

Stephanorhyssalus

Gen. et sp. nov

Valid

Belokobylskij & Jouault

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Braconidae. Genus includes new species S. longiscapus.

Tetragonula florilega

Sp. nov

Valid

Engel in Engel et al.

Miocene (Langhian)

Zhangpu amber

2021 In Paleoentomology  China

A stingless bee, a species of Tetragonula.

Thigmocosmus

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Heterogynaidae. Genus includes new species T. longicornis.

Tyrannomecia

Gen. et sp. nov

In press

Jouault & Nel

Paleocene

Menat Formation

2021 In Paleoentomology  France

An ant belonging to the stem group of the subfamily Myrmeciinae. Genus includes new species T. inopinata.

Vespula? hassiaca

Sp. nov

In press

Abels & Wedmann

Eocene

Messel pit

2021 In Paleoentomology  Germany

A member of the family Vespidae.

Xeris dorbnikensis

Sp. nov

Valid

Manukyan in Manukyan & Smirnova

Eocene

Baltic amber

Europe (Baltic Sea region)

A horntail. Originally described as a species of Xeris; Archibald & Rasnitsyn (2022) transferred it to the genus Urocerus.

Zigrasimecia goldingot

Sp. nov

In press

Zhuang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

An ant in the tribe Sphecomyrmini.

Hymenopteran Research


Clade Neuropterida

Neuropterans

Name Novelty Status Authors Age Type locality Country Notes Images

Ankyloleon

Gen. et sp. nov

Valid

Badano, Haug & Cerretti in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

An early representative of the clade Myrmeleontiformia. The type species is A. caudatus.

Astioberotha coutreti

Sp. nov

In press

Jouault

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Rhachiberothidae belonging to the subfamily Paraberothinae.

Burmotachinymphes

Gen. et sp. nov

Valid

Cao, Wang & Liu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Mesochrysopidae. Genus includes new species M. bilobata.

Calobabinskaia

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Babinskaiidae. The type species is C. xiai.

Fuscopolystoechotes

Gen. et sp. nov

Valid

Xu, Zheng, Shih & Wang in Xu et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Ithonidae. Genus includes new species F. reticulatus.

Karenina cuneiformis

Sp. nov

Valid

Nakamine

Early Cretaceous (Aptian)

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Mesochrysopidae.

Lichenipolystoechotes fenestratus

Sp. nov

Valid

Xu, Fang, Shih & Wang in Xu et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Ithonidae.

Osmylidia donnae

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Quilchena site

2021 In Paleoentomology  Canada
2021 In Paleoentomology  British Columbia

A protosmyline Osmylid lance lacewing

Osmylidia glastrai

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Klondike Mountain Formation

2021 In Paleoentomology  United States
2021 In Paleoentomology  Washington

A protosmyline Osmylid lance lacewing

Osmylidia picta

Comb. nov

Valid

(Hagen)

Priabonian

Baltic amber

2021 In Paleoentomology  Europe

A protosmyline Osmylid lance lacewing
Moved from "Protosmylus" picta (1913)

Osmylidia taliae

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Green River Formation
Parachute Creek Member

2021 In Paleoentomology  United States
2021 In Paleoentomology  Colorado

A protosmyline Osmylid lance lacewing

Osmyloberotha

Gen. et sp. nov

Valid

Khramov

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A beaded lacewing. Genus includes new species O. simpla.

Paradoxoberotha

Gen. et sp. nov

In press

Nakamine et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Rhachiberothidae. Genus includes new species P. chimaera.

Paraneliana

Gen. et sp. nov

Valid

Jouault & Nel

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A member of the family Babinskaiidae. Genus includes new species P. sennlaubi.

Rasnitsyneura

Gen. et comb. nov

Valid

Makarkin, Ansorge & Khramov

Early Jurassic (Toarcian)

2021 In Paleoentomology  Germany

A member of the family Ithonidae sensu lato belonging to the subfamily Epigambriinae. The type species is "Prohemerobius" aequabilis Bode (1953).

Sinonothochrysa

Gen. et sp. nov

In press

Huang & Nel in Huang et al.

Paleocene

Buxin Formation

2021 In Paleoentomology  China

A member of the family Chrysopidae. Genus includes new species S. zhangi

Stenobabinskaia

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Babinskaiidae. The type species is S. punctata.

Xiaobabinskaia

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Babinskaiidae. The type species is X. lepidotricha.

Neuropteran research

  • Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.
  • New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug et al. (2021), who also review the morphological diversity of fossil lacewing larvae.

Raphidiopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Archiinocellia protomaculata

comb nov

valid

(Engel)

Lutetian

Green River Formation

2021 In Paleoentomology  USA
2021 In Paleoentomology  Colorado

A raphidiid snakefly.
Moved from Archiinocellia protomaculata Engel (2011)

2021 In Paleoentomology 
Archiinocellia protomaculata

Electrobaissoptera liui

Sp. nov

In press

Jouault, Engel & Nel

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A baissopterid snakefly.

Megaraphidia antiquissima

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Tranquille Formation

2021 In Paleoentomology  Canada
2021 In Paleoentomology  British Columbia

A raphidiid snakefly.

Megaraphidia hopkinsi

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Allenby Formation

2021 In Paleoentomology  Canada
2021 In Paleoentomology  British Columbia

A raphidiid snakefly.

Megaraphidia klondika

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Klondike Mountain Formation

2021 In Paleoentomology  United States
2021 In Paleoentomology  Washington

A raphidiid snakefly.

Megaraphidia ootsa

Sp. nov

Valid

Archibald & Makarkin

Ypresian

"Unnamed unit" Ootsa Group

2021 In Paleoentomology  Canada
2021 In Paleoentomology  British Columbia

A raphidiid snakefly.

Mesoraphidia baekthoensis

Sp. nov

In press

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

2021 In Paleoentomology  North Korea

A member of the family Mesoraphidiidae.

Mesoraphidia ryonsangensis

Sp. nov

In press

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

2021 In Paleoentomology  North Korea

A member of the family Mesoraphidiidae.

Clade †Palaeodictyopteroidea

†Megasecoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Anhuihymen

Gen. et sp. nov

In press

Huang & Nel in Huang et al.

Middle Permian

Yinping Formation

2021 In Paleoentomology  China

A protohymenid Megasecoptera.
The type species is A. medianelongata.

†Palaeodictyoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Dictyoneura goujonorum

Sp. nov

In press

Logghe et al.

Carboniferous-Permian (Gzhelian-Asselian)

2021 In Paleoentomology  France

A member of Palaeodictyoptera.

Sandiella herbigi

Sp. nov

In press

Brauckmann, Herd & Leipner

Carboniferous (Moscovian)

2021 In Paleoentomology  Germany

A member of Palaeodictyoptera belonging to the family Eugereonidae.

Palaeodictyopteran research

  • A study on the anatomy of fossil specimens described as consecutive series of different larval stages of palaeodictyopteran species Tchirkovaea guttata and Paimbia fenestrata, evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).

Clade Palaeoptera

Ephemeroptera

Name Novelty Status Authors Age Type locality Country Notes Images

Astraeoptera

Gen. et sp. nov

Valid

Brandão et al.

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A mayfly of uncertain phylogenetic placement.
Genus includes new species A. cretacica.

Burmella

Gen. et 2 sp. nov

Valid

Godunko, Martynov & Staniczek

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A vietnamellid mayfly.
The type species is B. paucivenosa; genus also includes B. clypeata.

Protoligoneuria heloisae

Sp. nov

In press

Storari et al.

Early Cretaceous (Aptian)

Crato Formation

2021 In Paleoentomology  Brazil

A hexagenitid mayfly.

Triassodotes rasnitsyni

Sp. nov

Valid

Sinitshenkova

Triassic

2021 In Paleoentomology  Russia

A misthodotid mayfly.

Ephemeropteran research

  • Sroka et al. (2021) provide new information on the morphology of Permian stem-mayflies Misthodotes sharovi and M. zalesskyi from the Tshekarda locality (Kungurian Koshelevka Formation; Perm Krai, Russia), and infer the life history traits of both the adult and larval stages of these insects.
  • Possible underwater mayfly burrows are described from the Turonian Ferron Sandstone (Utah, United States) by King et al. (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new ichnospecies Glossifungites gingrasi.

Odonatoptera

Odonata

Name Novelty Status Authors Age Type locality Country Notes Images

Chalcolestes tibetensis

Sp. nov

In press

Xia et al.

Late Eocene

Nima Basin

2021 In Paleoentomology  China

A species of Chalcolestes.

Cretaphylolestes

Gen. et sp. nov

In press

Huang et al.

Early Cretaceous (Aptian)

Shouchang Formation

2021 In Paleoentomology  China

A damselfly belonging to the family Synlestidae. Genus includes new species C. cretacicus.

Dakotaeschnidium

Gen. et sp. nov

Valid

Nel

Late Cretaceous (Maastrichtian)

Fox Hills Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  South Dakota)

A dragonfly belonging to the family Aeschnidiidae. Genus includes new species D. spedeni.

Dysagrion pruettae

Sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Washington)

A member of the family Dysagrionidae.

2021 In Paleoentomology 
Dysagrion pruettae

Dysagrionites

Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Washington)

A collective group name for any fossil species referable to the Dysagrioninae whose orthotaxon generic affinity is unclear. Includes new species D. delinei.

Epilestes rasnitsyni

Sp. nov

Valid

Felker & Vasilenko

Permian

2021 In Paleoentomology  Russia

A damselfly belonging to the family Permagrionidae.

Jingguaeshna

Gen. et sp. nov

Valid

Zheng et al.

Miocene

2021 In Paleoentomology  China

A dragonfly belonging to the group Aeshnoidea, possibly a member of the family Telephlebiidae. Genus includes new species J. taoae.

Marado

Gen. et sp. nov

Valid

Petrulevičius

Paleocene

Maíz Gordo Formation

2021 In Paleoentomology  Argentina

A damselfly belonging to the family Coenagrionidae. The type species is M. marado.

Moltenophlebia

Gen. et sp. nov

Valid

Deregnaucourt et al.

Triassic

Molteno Formation

2021 In Paleoentomology  South Africa

A member of the stem group of Odonata, belonging to the group Zygophlebiida within the Triadophlebiomorpha. Genus includes new species M. lindae.

Okanagrion

Gen. et 8 sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Washington)

A member of the family Dysagrionidae. The type species is O. threadgillae; genus also includes new species O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum and O. worleyae.

2021 In Paleoentomology 
Okanagrion threadgillae

Okanopteryx

Gen. et 3 sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Tranquille Formation

2021 In Paleoentomology  Canada
(2021 In Paleoentomology  British Columbia)

A member of the family Dysagrionidae. The type species is O. macabeensis; genus also includes new species O. fraseri and O. jeppesenorum.

Phyonganpodagrion

Gen. et sp. nov

In press

So & Won

Early Cretaceous (BarremianAptian)

Sinuiju Formation

2021 In Paleoentomology  North Korea

A damselfly in the family Megapodagrionidae. Genus includes new species P. ryonsangae.

Protohemiphlebia

Fam. et gen. et 2 sp. nov

In press

Zheng, Jarzembowski & Nel in Zheng et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A stem hemiphlebioid damselfly belonging to the new family Protohemiphlebiidae. Genus includes new species P. zhangi and P. meiyingae.

Pulchrairina

Gen. et sp. nov

In press

Martynov, Vasilenko & Perkovsky

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A damselfly belonging to the group Coenagrionoidea.
Genus includes new species P. electra.

Republica

Gen. et sp. nov

Valid

Archibald & Cannings

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Washington)

A euphaeid damselfly.
Type species is R. weatbrooki.
Not to be confused with the plant genus Republica also from the formation.

2021 In Paleoentomology 
Republica weatbrooki

Stenodiafanus

Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States
(2021 In Paleoentomology  Washington)

A dysagrionid odonate.
Type species is S. westersidei.

2021 In Paleoentomology 
Stenodiafanus westersidei

Stenophlebia ryonsangensis

Sp. nov

In press

Won, So & Kim

Early Cretaceous

Sinuiju Formation

2021 In Paleoentomology  North Korea

A stenophlebiid odonate.

Whetwhetaksa

Fam. Nov., Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

2021 In Paleoentomology  United States

The type genus of Whetwhetaksidae, a new family of uncertain placement.
The type species is W. millerae.

2021 In Paleoentomology 
Whetwhetaksa millerae
Odonate research
  • Archibald et al. (2021) establish a new, extinct suborder of Odonata named Cephalozygoptera, close to but distinct from damselflies, composed of the families Dysagrionidae, Sieblosiidae and possibly Whetwhetaksidae; their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald et al. to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.

†Protozygoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Kennedya kedrovkensis

Sp. nov

Valid

Felker

Early Triassic

2021 In Paleoentomology  Russia

A member of Protozygoptera belonging to the family Kennedyidae.

Kennedya suchonensis

Sp. nov

Valid

Felker

Late Permian

2021 In Paleoentomology  Russia

Progoneura kemerovensis

Sp. nov

Valid

Felker

Early Triassic

2021 In Paleoentomology  Russia

A member of Protozygoptera belonging to the family Kennedyidae.

Other odonatopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Brunellopteron

Gen. et sp. nov

Valid

Béthoux, Deregnaucourt & Norrad in Béthoux et al.

Carboniferous (Moscovian)

Sunbury Creek Formation

2021 In Paleoentomology  Canada
(2021 In Paleoentomology  New Brunswick)

A member of Neodonatoptera of uncertain phylogenetic placement. The type species is B. norradi.

Osnabruggiala

Gen. et sp. nov

Valid

Zessin, Brauckmann & Gröning

Carboniferous (Pennsylvanian)

Osnabrück Formation

2021 In Paleoentomology  Germany

Probably a basal member of Odonatoptera. The type species is O. seppelti.

Clade †Paoliidea

†Paoliida

Name Novelty Status Authors Age Type locality Country Notes Images

Avionblattinopsis

Gen. et sp. nov

In press

Quispe et al.

Carboniferous (Moscovian)

2021 In Paleoentomology  France

A blattinopsid Paoliidan.
The type species is A. oudardi.

Blattinopsis indefinitus

Sp. nov

Valid

Rasnitsyn & Aristov

Permian (Kungurian)

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Krasnoyarsk Krai)

A Blattinopsidae Paoliidan

Glaphyrophlebia borea

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Komi Republic)

A Blattinopsid.

Glaphyrophlebia iva

Sp. nov

Valid

Aristov & Rasnitsyn

Middle Permian

2021 In Paleoentomology  Russia

A member of the family Blattinopsidae.

Glaphyrophlebia komia

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Komi Republic)

A member of the family Blattinopsidae.

Glaphyrophlebia vorkutensis

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Komi Republic)

A member of the family Blattinopsidae.

Clade Paraneoptera

Hemipterans

Name Novelty Status Authors Age Type locality Country Notes Images

Acrotiara

Trib. et gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Cixiidae; the type genus of the new tribe Acrotiarini. The type species is A. multigranulata Luo & Bourgoin.

Aneurus damzeni

Sp. nov

Valid

Heiss in Heiss, Kairišs & Bukejs

Eocene

Baltic amber

2021 In Paleoentomology  Russia
(2021 In Paleoentomology  Kaliningrad Oblast)

A species of Aneurus.

Aphelicophontes

Gen. et sp. nov

Valid

Swanson et al.

Eocene

Green River Formation

2021 In Paleoentomology  United States

A member of the family Reduviidae belonging to the subfamily Harpactorinae. Genus includes new species A. danjuddi.

Baltiomiris

Gen. et sp. nov

Valid

Kim, Chérot & Jung

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Miridae. The type species is B. herczeki.

Barremixius

Gen. et comb. nov

In press

Bourgoin & Szwedo in Luo et al.

Early Cretaceous

Weald Clay

2021 In Paleoentomology  United Kingdom

A member of the family Cixiidae; a new genus for "Cixius" petrinus Fennah.

Beaconiella cincta

Sp. nov

Valid

Shcherbakov

Triassic (LadinianCarnian)

Madygen Formation

2021 In Paleoentomology  Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.

Beaconiella pulchra

Sp. nov

Valid

Shcherbakov

Triassic (Ladinian–Carnian)

Madygen Formation

2021 In Paleoentomology  Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.

Burmacader bicoloripennis

Sp. nov

Valid

Souma, Yamamoto & Takahashi

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tingidae.

Burmacader grandis

Sp. nov

Valid

Heiss & Guilbert

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tingidae.

Burmala

Gen. et sp. nov

Valid

Liu et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Malmopsyllidae. Genus includes new species B. liaoyaoi.

Burmaprosbole

Gen. et sp. nov

Valid

Qiao et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tettigarctidae. Genus includes new species B. nervosa.

Burmavianaida

Gen. et sp. nov

Valid

Souma, Yamamoto & Takahashi

Cretaceous (Albian or Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tingidae. The type species is B. anomalocapitata.

Cretamystilus

Gen. et sp. nov

Valid

Kim, Lim & Jung

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Miridae belonging to the tribe Mecistoscelini. Genus includes new species C. herczeki.

Cretodorus rostellatus

Sp. nov

In press

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Mimarachnidae.

Cretomultinervis

Gen. et sp. nov

Valid

Fu & Huang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Sinoalidae. Genus includes new species C. burmensis.

Dachibangus hui

Sp. nov

Valid

Zhang, Yao & Pang in Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Mimarachnidae. Originally described as a species of Dachibangus, but subsequently transferred to the genus Xiaochibangus.

Delphitiara

Gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is D. tibiocoronata Luo & Bourgoin.

Dorytocus jiaxiaoae

Sp. nov

Valid

Song, Szwedo & Bourgoin in Song et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A planthopper.

Duyana

Gen. et sp. nov

Valid

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A leafhopper belonging to the subfamily Ledrinae. Genus includes new species D. electronica.

Electromyiomma herczeki

Sp. nov

Valid

Kim & Jung

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Miridae.

Eospinosus

Gen. et 2 sp. nov

Valid

Wedmann et al.

Eocene

Messel pit

2021 In Paleoentomology  Germany
2021 In Paleoentomology  United States
(2021 In Paleoentomology  Colorado)

A member of the family Pentatomidae. The type species is E. peterkulkai; genus also includes E. greenriverensis.

Ferriantenna

Gen. et sp. nov

Valid

Cumming & Le Tirant

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Coreidae belonging to the subfamily Coreinae. The type species is F. excalibur.

Hallex martinsnetoi

Sp. nov

In press

Monferran et al.

Early Cretaceous

Crato Formation

2021 In Paleoentomology  Brazil

A leafhopper.

Hypsotsijilia

Gen. et sp. nov

Valid

Cifuentes-Ruiz & Brailovsky in Cifuentes-Ruiz et al.

Early Miocene

La Quinta Formation
(Mexican amber)

2021 In Paleoentomology  Mexico

A member of the family Schizopteridae. The type species is H. bretoni.

Inoderbe

Fam. et gen. et sp. nov

Valid

Shcherbakov & Emeljanov

Cretaceous (Albian–Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A planthopper belonging to the new family Inoderbidae. The type species is I. rapunzel.

Juroala pulchra

Sp. nov

Valid

Fu & Huang

Late Jurassic

Daohugou Beds

2021 In Paleoentomology  China

A member of the family Sinoalidae.

Karatauella

Nom. nov

Valid

Doweld

Jurassic (Bathonian-Oxfordian)

2021 In Paleoentomology  Kazakhstan

A member of the family Protopsyllidiidae; a replacement name for Carpenterella Becker-Migdisova (1968).

Latidorsum

Gen. et sp. nov

Wang, Tang & Yao in Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tingidae. The type species is L. carinbifarium.

Microtingis

Gen. et sp. nov

Junior synonym

Heiss & Golub

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Tingidae. Genus includes new species M. leptosoma. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of Burmavianaida anomalocapitata.

Multistria

Gen. et sp. nov

Valid

Zhang, Yao & Pang in Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Mimarachnidae. The type species is M. orthotropa.

Paleocader balticus

Sp. nov

Valid

Heiss & Golub

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Tingidae.

Paleocader pulchellus

Sp. nov

Valid

Heiss & Golub

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Tingidae.

Paleocader rovnensis

Sp. nov

Valid

Heiss & Golub

Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Tingidae.

Parasinalda sukachevae

Sp. nov

Valid

Golub, Perkovsky & Vasilenko

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A member of the family Tingidae.

Pentacarinus

Gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is P. kachinensis Luo & Bourgoin.

Pseudocaulisoculus

Gen. et sp. nov

Valid

Kóbor & Roca-Cusachs

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Yuripopovinidae. Genus includes new species P. longicornis.

Pubivetanthocoris

Gen. et sp. nov

In press

Tang, Wang & Yao in Tang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Vetanthocoridae. Genus includes new species P. carinalis.

"Pyrrhocoris" rottensis

Nom. nov

Valid

Kment

Late Oligocene

2021 In Paleoentomology  Germany

A member of Lygaeoidea of uncertain phylogenetic placement; a replacement name for Pyrrhocoris tibialis Statz & Wagner (1950).

Rovnoxestus

Gen. et sp. nov

Valid

Dietrich, Dmitriev & Perkovsky

Late Eocene

Rovno amber

2021 In Paleoentomology  Ukraine

A leafhopper. Genus includes new species R. rasnitsyni.

Saltissus fennahi

Sp. nov

In press

Luo, Liu & Jarzembowski in Luo et al.

Early Cretaceous

Weald Clay

2021 In Paleoentomology  United Kingdom

A member of the family Mimarachnidae.

Scytinoptera tongchuanensis

Sp. nov

In press

Zhang, Zheng & Zhang in Zhang et al.

Middle Triassic

Tongchuan Formation

2021 In Paleoentomology  China

A member of the family Scytinopteridae.

Sinogranulus

Gen. et sp. nov

In press

Zhang, Chen & Zhang

Middle Triassic

Tongchuan Formation

2021 In Paleoentomology  China

A member of Scytinopteroidea belonging to the family Granulidae. Genus includes new species S. qishuiheensis.

Spinonympha

Gen. et sp. nov

Valid

Luo, Wang & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A planthopper of uncertain phylogenetic placement. The type species is S. shcherbakovi.

Szeiinia

Fam. nov., Gen. et sp. nov

Valid

Zhang et al.

Late Triassic

Yanchang Formation

2021 In Paleoentomology  China

A planthopper in the new family Szeiiniidae. Genus includes new species S. huanglongensis

Tennentsia orientalis

Sp. nov

In press

Fu, Azar & Huang

Middle Triassic

Yanchang Formation

2021 In Paleoentomology  China

A member of Cicadomorpha belonging to the family Dysmorphoptilidae.

Triknightia

Gen. et sp. nov

Valid

Shcherbakov

Triassic (Ladinian–Carnian)

Madygen Formation

2021 In Paleoentomology  Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Scytinopteroidea and the family Paraknightiidae. The type species is T. mira.

Tuthillia maculosa

Sp. nov

Valid

Poinar & Brown

Dominican amber

2021 In Paleoentomology  Dominican Republic

A member of Psylloidea.

Valdicopis

Gen. et sp. nov

Valid

Li, Chen & Jarzembowski

Early Cretaceous

2021 In Paleoentomology  United Kingdom

A member of the family Procercopidae. Genus includes new species V. tonyi.

Youngeewa simillima

Sp. nov

Valid

Gębicki & Walczak in Gębicki, Walczak & Zmarzły

Eocene

Baltic amber

Europe (Baltic Sea region)

A leafhopper belonging to the subfamily Mileewinae.

Hemipteran research

  • Grimaldi & Vea (2021) reinterpret Mesophthirus engeli as a scale insect, and reject the interpretation of this insect as an ectoparasite.
  • Redescription of the holotype specimen of Paranoika placida is published by Moura-Júnior et al. (2021), who transfer this species to the genus Lethocerus, and consider this species to be the senior synonym of Lethocerus vetus.
  • A fulgorid specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu et al. (2021).
  • Revision of the fossil record of Fulgoromorpha and Cicadomorpha from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).

Psocodeans

Name Novelty Status Authors Age Type locality Country Notes Images

Burmesopsocus

Gen. et sp. nov

Valid

Yoshizawa in Yoshizawa & Yamamoto

Cretaceous (Albian–Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Homilopsocidea of uncertain phylogenetic placement. The type species is B. lienhardi.

Cormopsocus baleoi

Sp. nov

Valid

Hakim, Azar & Huang

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Trogiomorpha belonging to the family Cormopsocidae.

Cormopsocus neli

Sp. nov

Valid

Hakim, Azar & Huang in Hakim et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Trogiomorpha belonging to the family Cormopsocidae.

Empherium

Gen. et sp. nov

Valid

Hakim, Huang & Azar

Late Cretaceous

2021 In Paleoentomology  Russia

A member of Trogiomorpha belonging to the family Empheriidae. Genus includes new species E. rasnitsyni.

Eolepinotus zherikhini

Sp. nov

Valid

Hakim, Huang & Azar

Late Cretaceous

2021 In Paleoentomology  Russia

A member of Trogiomorpha belonging to the family Trogiidae.

Heliadesdakruon

Gen. et sp. nov

Valid

Cumming & Le Tirant

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Trogiomorpha belonging to the family Archaeatropidae. The type species is H. morganae.

Longiglabellus

Gen. et 2 sp. nov

In press

Wang, Li & Yao in Wang et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Cormopsocidae. Genus includes new species L. edentatus and L. pedhyalinus.

Palaeosiamoglaris hammanaensis

Sp. nov

In press

Hakim, Huang & Azar

Early Cretaceous (Barremian)

Lebanese amber

2021 In Paleoentomology  Lebanon

A member of Psocodea belonging to the family Prionoglarididae.

Palaeosiamoglaris hkamtiensis

Sp. nov

In press

Jouault et al.

Early Cretaceous

Hkamti amber

2021 In Paleoentomology  Myanmar

A member of Psocodea belonging to the family Prionoglarididae.

Psyllipsocus myanmarensis

Sp. nov

In press

Jouault et al.

Late Cretaceous (Cenomanian)

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.

Psyllipsocus yangi

Sp. nov

Valid

Liang & Liu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.

Stimulopsocus

Gen. et sp. nov

In press

Liang & Liu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A member of the family Cormopsocidae. Genus includes new species S. jiewenae.

Psocodean research

  • Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller et al. (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.

Clade Perlidea

Dermapterans

Name Novelty Status Authors Age Type locality Country Notes Images

Aglyptoderma

Gen. et 2 sp. nov

Valid

Xiong et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A semenoviolid eodermapteran earwig.
Genus includes the new species A. elongatum and A. cylindratum.

Aneuroderma

Gen. et sp. nov

Valid

Xiong, Engel & Ren in Xiong et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Protodiplatyidae. The type species is A. oiodes.

Dacryoderma

Gen. et comb. nov

Valid

Engel

Early Jurassic (Sinemurian)

Charmouth Mudstone Formation

2021 In Paleoentomology  United Kingdom

A dermapterid earwig.
The type species is "Brevicula" teres Tihelka (2019).

Eminepygia

Gen. et sp. nov

Valid

Chen & Zhang

Cenomanian

Burmese amber

2021 In Paleoentomology  Myanmar

A pygidicranid earwig.
Genus includes new species E. myanmarensis.

Sinopalaeodermata concavum

Sp. nov

Valid

Xiong, Engel & Ren in Xiong et al.

Middle Jurassic

Jiulongshan Formation

2021 In Paleoentomology  China

A member of the family Dermapteridae.

Plecoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Balticonemoura

Gen. et sp. nov

In press

Chen

Eocene

Baltic amber

Europe (Baltic Sea region)

A nemourid stonefly.
Genus includes new species B. bulbosus.

Billoperla

Gen. et sp. nov

In press

Chen & Xu

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A perlid stonefly belonging to the family.
Genus includes new species B. starki.

Isoperla baltica

Sp. nov

Valid

Jouault et al.

Eocene

Baltic amber

Europe (Baltic Sea regon)

A perlodid stonefly.

Other insects

Name Novelty Status Authors Age Type locality Country Notes Images

Anakitala

Gen. et sp. nov

Valid

Aristov

Late Permian

2021 In Paleoentomology  Russia

A mesorthopterid Eoblattidan Polyneopteran.
The type species is A. intermittendia.

Baharellinus elegans

Sp. nov

Valid

Aristov

Late Permian

2021 In Paleoentomology  Russia

A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae.

Belmophenopterum rasnitsyni

Sp. nov

Valid

Storozhenko & Aristov

Triassic (Ladinian-Carnian)

Madygen Formation

2021 In Paleoentomology  Kyrgyzstan

A member of the family Mesorthopteridae.

Cabarzopterum

Gen. et sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

2021 In Paleoentomology  Germany

A member of Grylloblattida of uncertain phylogenetic placement. Genus includes new species C. magnificus.

Delopterinus alsenzensis

Sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

2021 In Paleoentomology  Germany

A miomopteran belonging to the family Palaeomanteidae.

Delopterum novokshonovi

Sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

2021 In Paleoentomology  Germany

A miomopteran belonging to the family Palaeomanteidae.

Elmocladus

Gen. et comb. nov

Valid

Rasnitsyn & Aristov

Early Permian

2021 In Paleoentomology  United States

A member of Polyneoptera belonging to the group Eoblattida and the family Tococladidae. The type species is "Opisthocladus" strictus Carpenter (1976).

Galloembia

Gen. et sp. nov

Valid

Falières, Engel & Nel

Earliest Eocene

Oise amber

2021 In Paleoentomology  France

A webspinner belonging to the family Embiidae. The type species is G. raholai.

Gnethoda odontophora

Sp. nov

In press

Lai et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A webspinner belonging to the family Clothodidae.

Kamopanorpa rasnitsyni

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

2021 In Paleoentomology  Russia

A member of the (stem-trichopteran or stem-amphiesmenopteran) family Microptysmatidae.

Kamopanorpa rotunda

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

2021 In Paleoentomology  Russia

A member of the family Microptysmatidae.

Kamopanorpa sivchikovi

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

2021 In Paleoentomology  Russia

A member of the family Microptysmatidae.

Liomopterites sperbersbachensis

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

2021 In Paleoentomology  Germany

A member of Grylloblattida belonging to the family Liomopteridae.

Liomopterum fuscatum

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

2021 In Paleoentomology  Germany

A member of Grylloblattida belonging to the family Liomopteridae.

Madygembia

Gen. et sp. nov

Valid

Aristov & Storozhenko

Triassic (Ladinian or Carnian)

Madygen Formation

2021 In Paleoentomology  Kyrgyzstan

A webspinner belonging to the family Alexarasniidae. The type species is M. rasnitsyni.

Magicivena

Fam. et gen. et 3 sp. nov

Valid

Yang et al.

Early Cretaceous

Yixian Formation

2021 In Paleoentomology  China

A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family Magicivenidae. The type species is M. sticta; genus also includes M. elegans and M. antennalis.

Multivena

Gen. et sp. nov

In press

Lai et al.

Cretaceous

Burmese amber

2021 In Paleoentomology  Myanmar

A webspinner belonging to the family Sorellembiidae. Genus includes new species M. curvivena.

Pictoborella

Gen. et sp. et comb. nov

Valid

Cui et al.

Early Permian

2021 In Paleoentomology  Germany

A member of Grylloblattida. Genus includes new species P. clara, as well as "Oborella" germanica Prokop et al. (2021).

Rasnalexia

Fam. et gen. et sp. nov

Valid

Gorochov

Triassic

2021 In Paleoentomology  Kyrgyzstan

A member of Embioptera belonging to the new family Rasnalexiidae. Genus includes new species R. rasnitsyni.

Uralioma thuringiensis

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

2021 In Paleoentomology  Germany

A member of Grylloblattida belonging to the family Liomopteridae.

Research

  • A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the Permian (Wordian) Pelitic Formation (France) by Logghe et al. (2021), who name a new ichnotaxon Phasmichnus radagasti.
  • A dragonfly-type odonatopteran larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).

General research

  • Revision of the fossil record of insects from the Triassic of China is published by Zhang et al. (2021).
  • A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).

References

Tags:

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