Hippoidea

Hippoidea is a superfamily of decapod crustaceans known as mole crabs or sand crabs.

Hippoidea Temporal range: Maastrichtian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Blepharipoda occidentalis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Malacostraca
Order:	Decapoda
Suborder:	Pleocyemata
Infraorder:	Anomura
Superfamily:	Hippoidea Latreille, 1825
Families
Albuneidae Blepharipodidae Hippidae

Hippoids are adapted to burrowing into sandy beaches, a habit they share with raninid crabs, and the parallel evolution of the two groups is striking. In the family Hippidae, the body is almost ovoid, the first pereiopods have no claws, and the telson is long, none of which are seen in related groups. Unlike most other decapods, sand crabs cannot walk; instead, they use their legs to dig into the sand. Members of the family Hippidae beat their uropods to swim.

Apart from the polar regions, hippoids can be found on beaches throughout the world. Larvae of one species have also been found in Antarctic waters, despite the lack of suitable sandy beaches in the Antarctic.

Alongside hermit crabs and allies (Paguroidea), squat lobsters and allies (Galatheoidea) and the hairy stone crab (Lomis hirta, Lomisoidea), Hippoidea is one of the four groups that make up the infraorder Anomura. Of the four, Hippoidea is thought to be the most basal, with the other three groups being more closely related to each other than to Hippoidea.

The fossil record of sand crabs is sparse, but extends back to the Cretaceous period. Sand crabs are placed in three families (exclusively fossil taxa are marked †):