Panorpida

Panorpida or Mecopterida is a proposed superorder of Holometabola. The conjectured monophyly of the Panorpida is historically based on morphological evidence, namely the reduction or loss of the ovipositor and several internal characteristics, including a muscle connecting a pleuron and the first axillary sclerite at the base of the wing, various features of the larval maxilla and labium, and basal fusion of CuP and A1 veins in the hind wings. The monophyly of the Panorpida is supported by recent molecular data.

Panorpida
Panorpida
Celastrina argiolus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Clade: Aparaglossata
Superorder: Panorpida
Clades
part of Holometabola

Hymenoptera (sawflies, wasps, ants, bees) Panorpida

Panorpida
Amphiesmenoptera

Trichoptera (caddisflies) Panorpida

Lepidoptera (butterflies and moths) Panorpida

Antliophora

Antliophora

The Panorpid clade Antliophora contains one of the major phylogenetic puzzles among the Insecta. It is unclear as of 2020 whether the Mecoptera (scorpionflies and allies) form a single clade, or whether the Siphonaptera (fleas) are inside that clade, so that the traditional "Mecoptera" is paraphyletic. However the earlier suggestion that the Siphonaptera are sister to the Boreidae (snow scorpionflies) is not supported; instead, there is the possibility that they are sister to another Mecopteran family, the Nannochoristidae of the Southern hemisphere. The two possible trees are shown below:

(a) Mecoptera is paraphyletic, containing Siphonaptera:

Antliophora

Diptera (true flies) Panorpida 

Pistillifera (scorpionflies, hangingflies, 400 spp.) Panorpida 

Nannochoristidae (southern scorpionflies, 8 spp.)

Siphonaptera (fleas, 2500 spp.) Panorpida 

Boreidae (snow scorpionflies, 30 spp.) Panorpida 

(b) Mecoptera is monophyletic, sister to Siphonaptera

Antliophora

Diptera (true flies) Panorpida 

Mecoptera

Pistillifera (scorpionflies, hangingflies, 400 spp.) Panorpida 

Boreidae (snow scorpionflies, 30 spp.) Panorpida 

Nannochoristidae (southern scorpionflies, 8 spp.)

Siphonaptera (fleas, 2500 spp.) Panorpida 


References

Further reading

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