Accouplement de Neorrhina punctatum .
Famille
Les Scarabaeidae , en français Scarabéidés , sont une famille d'insectes coléoptères de la super-famille des Scarabaeoidea. Ils sont connus sous le nom vernaculaire de Scarabée . C'est notamment la famille des dynastes, du scarabée rhinocéros européen, des bousiers et des hannetons.
Les limites de cette famille sont encore débattues et controversées. Les chercheurs européens en ont une définition plutôt restrictive, alors qu'un certain nombre de chercheurs américains y incorporent bien plus de taxons.
Les antennes des insectes de cette famille sont en massue dont l'extrémité peut s'ouvrir en un éventail de feuillets. L'extrémité de l'abdomen est découverte en raison d'élytres normalement tronqués. La plupart des espèces volent correctement. Plusieurs de ces espèces stridulent en frottant l'extrémité de leurs élytres contre la face dorsale de leur abdomen. Cette vaste famille comprend des coprophages et des saproxylophages nuisibles aux végétaux.
Noms vernaculaires
Le terme de Scarabée provient, via le vieux français « escarbot », du latin scarabaeus , par altération du grec ancien κάραβος / kárabos , « escarbot, scarabée, insecte », également à l'origine du terme carabe. Le terme « scarabée» désigne de façon générale en zoologie l'ensemble de la super-famille des Scarabéoïdes, mais ce terme peut désigner chez certains auteurs, d'une façon plus restrictive, la famille des Scarabaeidae, et même pour certains uniquement les coprophages de cette famille[réf. nécessaire] .
Plusieurs groupes, qui peuvent être paraphylétiques, de ces insectes ont des noms vernaculaires tels que les Dynastes ou scarabées rhinocéros, les bousiers, les hannetons... Certaines espèces ont un nom vernaculaire particulier comme le Pique-prune (Osmoderma eremita ).
Systématique
La famille a été décrite par l'entomologiste français Pierre André Latreille en 1802.
Liste des sous-familles Selon le Catalogue of Life (23 juin 2023) , cette famille se divise en 28 sous-familles :
Aclopinae Blanchard, 1850 Aegialiinae Castelnau, 1840 Allidiostomatinae Arrow, 1940 Aphodiinae Leach, 1815 Aulonocneminae Janssens, 1946 Cetoniinae Leach, 1815 Chironinae Blanchard, 1845 † Cretoscarabaeinae Nikolajev, 1995 Dynamopodinae Arrow, 1911 Dynastinae MacLeay, 1819 Eremazinae Stebnicka, 1977 Hopliinae Latreille, 1829 Lichniinae Burmeister, 1844 † Lithoscarabaeinae Nikolajev, 1992 Melolonthinae MacLeay, 1819 Oncerinae LeConte, 1861 Orphninae Erichson, 1847 Pachydeminae Reitter, 1902 Phaenomeridinae Erichson, 1847 Podolasiinae Howden, 1997 † Prototroginae Nikolajev, 2000 Rutelinae MacLeay, 1819 Scarabaeinae Latreille, 1802 Sericinae Kirby, 1837 Sericoidinae Erichson, 1847 Termitotroginae Wasmann, 1918 Trichiinae Kolbe, 1897 Valginae Schenkling, 1922 Liste des genres Selon le Catalogue of Life (23 juin 2023) , cette famille regroupe 2 590 genres :
Ablabera Dejean, 1833 Ablaberoides Blanchard, 1850 Acanthaphodius Schmidt, 1909 Acanthobodilus Dellacasa, 1983 Acanthonitis Janssens, 1937 Acanthovalgus Kraatz, 1895 Aceratus Prell, 1936 Acheilo Britton, 1988 Achelyna Erichson, 1848 Achloa Erichson, 1840 Achranoxia Kraatz, 1888 Aclopus Erichson, 1835 Acoma Casey, 1889 Acraspedon Arrow, 1899 Acrobolbia Ohaus, 1912 Acrossidius Schmidt, 1914 Acrossoides Schmidt, 1914 Acrossus Mulsant, 1842 Acrothyrea Kraatz, 1882 Actinobolus Westwood, 1841 Acylochilus Ohaus, 1909 Adebrattia Bordat & Howden, 1995 Adelaeus Ratcliffe, 2014 Adeloparius Schmidt, 1914 Adoretops Kraatz, 1883 Adoretosoma Blanchard, 1851 Adoretus Dejean, 1833 Adorodocia Brenske, 1893 Adoroleptus Brenske, 1893 Adorrhinyptia Arrow, 1917 Adoryphorus Blackburn, 1889 Adossa Britton, 1995 Adraria Villiers, 1956 Aegialia Latreille, 1807 Aegidiellus Paulian, 1984 Aegidinus Arrow, 1904 Aegidium Westwood, 1845 Aegopsis Burmeister, 1847 Aegostheta Dejean, 1833 Aequatoria Arrow, 1899 Aethiaratrogus Decelle, 1982 Aethiessa Burmeister, 1842 Afroataenius Petrovitz, 1969 Afrodiapterna Dellacasa, 1984 Afrodiastictus Pittino & Mariani, 1986 Afrodrepanus Krikken, 2009 Afroemadiellus Bordat, 2019 Afrolepis Decelle, 1968 Afrotrichonotulus Bordat, 2016 Agamopus Bates, 1887 Aganocrossus Reitter, 1895 Agaocephala Le Peletier & Serville, 1828 Agaocnemis Moser, 1918 Agestrata Eschscholtz, 1829 Aglaphyra Brenske, 1896 Agnorimus Miyake & Iwase, 1991 Agoliinus Schmidt, 1914 Agolius Mulsant & Rey, 1869 Agrilinellus Dellacasa, Dellacasa & Gordon, 2008 Agrilinus Mulsant & Rey, 1869 Ahermodontus Báguena, 1930 Aidophus (Balthasar, 1963) Airapus Stebnicka & Howden, 1996 Ajmeraphodius Král, Mencl & Rakovič, 2021 Alaia Petrovitz, 1980 Alepida Allsopp, 2018 Aliaclitopa Lacroix, 2003 Alissonotum Arrow, 1908 Aliuscanthoniola Deschodt & Scholtz, 2008 Allara Britton, 1955 Alleucosma Schenkling, 1921 Allidiostoma Arrow, 1940 Alloblackburneus Bordat, 2009 Allobodilus Petrovitz, 1963 Allogymnopleurus Janssens, 1940 Allokotarsa Péringuey, 1904 Allomyrina Arrow, 1911 Allonitis Janssens, 1936 Allophileurinus Dupuis & Dechambre, 2001 Allophyllus Fauvel, 1903 Allorrhina Burmeister, 1842 Alloscelus Boucomont, 1923 Allothnonius Britton, 1978 Alocoderus Schmidt, 1914 Alogistotarsa Péringuey, 1904 Alvarengius Frey, 1975 Alvarinus Blanchard, 1850 Amadotrogus Reitter, 1902 Amaurina Kolbe, 1895 Amazula Kraatz, 1882 Amblochilus Blanchard, 1851 Amblonoxia Reitter, 1902 Amblymelanoplia Dombrow, 2002 Amblyodus Westwood, 1878 Amblyoproctus Kolbe, 1910 Amblyterus MacLeay, 1819 Amerilochus Skelley, 2007 Amerisaprus Stebnicka & Skelley, 2003 Amidorus Mulsant, 1870 Amietina Cambefort, 1981 Amiserica Nomura, 1974 Amithao Thomson, 1878 Ammoecioides Bordat, 1999 Ammoecius Mulsant, 1842 Amorphochelus Fairmaire, 1898 Amphiceratodon Huchet, 2000 Amphimallon Latreille, 1825 Amphistomus Lansberge, 1874 Amphitrichia Brenske, 1894 Ampliodactylus Smith, 2008 Ampotis Dechambre & Madge, 1979 Anacamptorrhina Blanchard, 1842 Anacanthodes Britton, 1990 Anaetius Petrovitz, 1969 Anagnathocera Arrow, 1922 Anahi Martinez, 1958 Analleucosma Antoine, 1989 Anamatochoranus Lacroix, 1995 Anartioschiza Kolbe, 1894 Anatista Brême, 1844 Anatona Burmeister, 1842 Anatonochilus Péringuey, 1907 Ancistrosoma Curtis, 1835 Ancognatha Erichson, 1847 Ancylonyx Britton, 1957 Anelaphinis Kolbe, 1892 Anenaria Lacroix, 1993 Aneucomides Blackburn, 1898 Anisocanthon Martinez & Pereira, 1956 Anisochelus Burmeister, 1844 Anisonyx Latreille, 1807 Anisopholis Moser, 1913 Anisoplia Fischer Von Waldheim, 1824 Anisorrhina Westwood, 1842 Ankaratrotrox Paulian, 1954 Annegialia Howden, 1971 Anochilia Burmeister, 1842 Anocoela Moser, 1914 Anoctus Sharp, 1875 Anodontopopillia Ohaus, 1901 Anomacaulus Fairmaire, 1878 Anomala Samouelle, 1819 † Anomalites Fric, 1885 Anomalochela Moser, 1913 Anomalomorpha Arrow, 1908 Anomalophylla Reitter, 1887 Anomalorhina Jameson, Paucar-Cabrera & Solis, 2003 Anomalurobius Paulian, 1952 Anomiopsoides Blackwelder, 1944 Anomiopus Westwood, 1842 Anomioserica Arrow, 1946 Anomius Mulsant & Rey, 1869 Anomolyna Fairmaire, 1897 Anomonyx Saylor, 1940 Anonychonitis Janssens, 1950 Anoplanomala Arrow, 1917 Anoplocheilus MacLeay, 1838 Anoplodrepanus Simonis, 1981 Anoplognatho Rivers, 1889 Anoplognathus Leach, 1815 Anoplostethus Brullé, 1837 Anoronotum Arrow, 1939 Anoxia Laporte, 1832 Anoxoides Petrovitz, 1971 Antandroyanus Lacroix, 1993 Antecessorirhyparus Minkina, 2020 † Antemnacrassa Gomez Pallerola, 1979 Anthoplia Medvedev, 1949 Anthotocus Britton, 1957 Anthracophora Burmeister, 1842 Anthracophorides Moser, 1918 Anthrencya Lacroix, 1993 Anticheira Eschscholtz, 1818 Anticheiroides Soula, 1998 † Antiqusolidus Lu, Ahrens, Bai, Shih & Ren, 2023 Antitrochalus Brenske, 1900 Antitrogus Burmeister, 1855 Antodon Brême, 1844 Aparammoecius Petrovitz, 1958 Apeltastes Howden, 1968 Aphaenoserica Brenske, 1900 Aphanesia Britton, 1990 Aphanesthes Kraatz, 1880 Aphelinis Antoine, 1987 Aphengium Harold, 1868 Aphengoecus Péringuey, 1901 Aphodaulacus Koshantschikov, 1911 Aphodiellus Schmidt, 1914 Aphodiopsis Paulian, 1942 Aphodius Hellwig, 1798 Aphodobius Péringuey, 1901 Aphodoharmogaster Endrödi, 1983 Aphodopsammobius Endrödi, 1964 Aphonides Rivers, 1889 Aphonodelus Kolbe, 1905 Aphonus LeConte, 1856 Aphotaenius Cartwright, 1952 Apicencya Lacroix, 1993 Aplopsis Blanchard, 1850 Apocnosoides Antoine, 2001 Apogonia Kirby, 1818 Aporecolpa Lansberge, 1886 Aposchiza Brenske, 1903 Apotolamprus Olsoufieff, 1947 Apotriodonta Baraud, 1962 Apsteiniella Schmidt, 1917 Aptenocanthon Matthews, 1974 Apterepilissus Montreuil, 2011 Apteroathlia Smith & Evans, 2018 Apterodemidea Gutiérrez, 1952 Aptychonitis Janssens, 1937 Araboplia Uliana & Sabatinelli, 2017 Arachnodes Westwood, 1842 Archedinus Morón & Krikken, 1990 Archeohomaloplia Nikolajev, 1982 † Archeopsammoporus Minkina, 2020 Archocamenta Brenske, 1895 Archophanes Kolbe, 1905 Archophileurus Kolbe, 1910 Archoserica Brenske, 1897 Arctodium Burmeister, 1844 Ardella Paulsen, 2021 Areoda MacLeay, 1819 Argentophileurus Penco & Zubarán, 2013 Argeremazus Stebnicka & Dellacasa, 2003 Argoplia Dombrow, 2003 Argyripa Thomson, 1878 Arielina Rossi, 1958 Arraphytarsa Péringuey, 1904 Arriguttia Martinez, 1960 Arupaia Stebnicka, 1999 Asactopholis Brenske, 1894 Asaphomorpha Brancsik, 1893 Aschnarhyparus Makhan, 2006 Asiactenius Nikolajev, 2000 Asiopertha (Medvedev, 1949) Asophrops Petrovitz, 1961 Aspidolea Bates, 1888 Aspilochilus Rojkoff, 2013 Aspilus Westwood, 1874 Astaena Erichson, 1847 Astaenoplia Martinez, 1957 Asthenopholis Brenske, 1898 Asthenorhella Westwood, 1874 Asthenorhina Westwood, 1843 Astibicola Britton, 1990 Astoxenus Péringuey, 1907 Astroscara Schürhoff, 1937 Ataeniopsis Petrovitz, 1973 Ataenius Harold, 1867 Atanyproctus Petrovitz, 1954 Ateuchetus Bedel, 1892 † Ateuchites Meunier, 1898 Ateuchus Weber, 1801 Athesphatoplia Lacroix, 1998 Athlia Erichson, 1835 Atlantemolanum González-Alvarado, Molano-Rendón & Vaz-de-Mello, 2019 Atrichelaphinis Kraatz, 1898 Atrichiana Distant, 1911 Attavicinus Philips & Bell, 2008 Atysilla Strand, 1942 Augoderia Burmeister, 1847 Augosoma Burmeister, 1841 Aulacopalpus Guérin-Méneville, 1838 Aulacopris White, 1859 Aulacoschiza Decelle, 1973 Aulacoserica Brenske, 1900 Aulonocnemis Klug, 1838 Aurum Hutchinson & Moeseneder, 2019 Ausmontins Deschodt & Davis, 2018 Australagolius Mencl, 2008 Australammoecius Petrovitz, 1958 Australaphodius (Balthasar, 1942) Australoxenella Howden & Storey, 1992 Automolius Britton, 1978 Autoserica Brenske, 1897 Axillonia Krikken, 2018 Bacchusia Mikšić, 1976 Badelina Thomson, 1880 Badiasis Machatschke, 1970 Badizoblax Thomson, 1877 † Baisarabaeus Nikolajev, 2005 Balanogonia Paucar-cabrera, 2003 Balbera Fairmaire, 1898 Ballucus Gordon & Skelley, 2007 Baloghonthobium Paulian, 1986 Balsameda Thomson, 1880 Barryfilius Allsopp, 2022 Barutus Ratcliffe, 1981 Barybas Blanchard, 1850 Basilewskyanus Endrödi, 1956 Basilewskynia Schein, 1957 Batesiana Chalumeau, 1983 Bdelyropsis Pereira & Vulcano & Martinez, 1960 Bdelyrus Harold, 1869 Beckhoplia Dombrow, 2005 Beriqua Péringuey, 1904 Besourenga Vaz-de-Mello, 2008 Bietia Fairmaire, 1898 Bilga Fairmaire, 1893 Bilobatus Machatschke, 1970 Biphyllocera White, 1841 Biralus Mulsant & Rey, 1869 Bisallardiana Antoine, 2003 Bisencya Lacroix, 1993 Bivalgus Paulian, 1961 Bizanus Péringuey, 1902 Blabephorus Fairmaire, 1898 Blackburneus Schmidt, 1914 Blaesia Burmeister, 1842 Blanchardoplia Lacroix, 1998 Blebea Lacroix, 1994 Blepharotoma Blanchard, 1850 Blikana Péringuey, 1902 Blitopertha Reitter, 1903 Bobiricola Endrödi, 1973 Bodiloides Dellacasa & Dellacasa, 2005 Bodilopsis Adam, 1994 Bodilus Mulsant & Rey, 1869 Bohepilissus Paulian, 1975 Bolax Fisher Von Waldheim, 1829 Bolbites Harold, 1868 Boletoscapter Matthews, 1974 Bombodes Westwood, 1848 Bonoraella Ruter, 1978 Bordatius Pittino & Mariani, 1986 Boreocanthon Halffter, 1958 Bosphorus Şenyüz, 2017 Bothrorrhina Burmeister, 1842 Bothynus Hope, 1837 Boucomontiellus Balthasar, 1932 Brachagenius Kraatz, 1890 Brachiaphodius Koshantschikov, 1913 Brachylepis Kolbe, 1894 Brachyllus Brenske, 1896 Brachymacroma Kraatz, 1896 Brachymis Thomson, 1858 Brachypholis Brenske, 1898 Brachysiderus Waterhouse, 1881 Brachysternus Guérin-Méneville, 1831 Bradypodidium Vaz-de-Mello, 2008 Brahmina Blanchard, 1851 Brancoplia Baraud, 1986 Brenskeiella Berg, 1898 Breviclypeus Ricchiardi, 2017 Bricoptis Burmeister, 1842 Brindalus Landin, 1960 Brittonius Özdikmen & Demir, 2008 Bruchaphodius Martinez, 1952 Bruneixenus Howden & Storey, 1992 Bubas Dejan, 1833 Buettikeria Sabatinelli & Pontuale, 1998 Bunbunius Nomura, 1970 Burmeisteriellus Berg, 1898 Burmeistoplia Dombrow, 2003 Butozania Mikšić, 1955 Byrasba Harold, 1869 Byrrhidium Harold, 1869 Byrrhomorpha Blackburn, 1892 Byrsalepis Brenske, 1898 Byrsopolis Burmeister, 1844 Caccobiomorphus Balthasar, 1964 Caccobius Thomson, 1859 Caeconthobium Paulian, 1984 Caelius Lewis, 1895 Caelorrhina Hope, 1841 Caesariatoplia Lacroix, 1998 Calamosternus Motschulsky, 1859 † Calcitoryctes Krell, 2011 Calicnemis Laporte, 1832 Caligodorus Gordon & Skelley, 2007 Callabonica Blackburn, 1895 Calliferoplia Lacroix, 1998 Callipechis Burmeister, 1842 Callirhinus Blanchard, 1851 Callistemonus Péringuey, 1901 Callistethus Blanchard, 1851 Callistopopillia Ohaus, 1903 Calloodes White, 1845 Callophylla Moser, 1916 Calloserica Brenske, 1894 Callynomes Mohnike, 1873 Calocolobopterus Dellacasa, 1984 Calodactylus Blanchard, 1850 Calomacraspis Bates, 1888 Calometopidius Bourgoin, 1917 Calometopus Blanchard, 1850 Calypsoryctes Howden, 1970 Camapterus Ricchiardi, 2000 Cambefortantus Paulian, 1986 Cambefortius Branco, 1989 Camenta Erichson, 1847 Camentoserica Brenske, 1900 Camerounophylla Lacroix, 2005 Camerunopholis Lacroix, 2002 Campsiura Hope, 1831 Campulipus Kirby, 1827 Campylophyllus Decelle, 1979 Campylotrochalus Brenske, 1900 Candezeollus Stebnicka & Howden, 1995 Canestera Saylor, 1938 Canthidium Erichson, 1847 Canthochilum Chapin, 1934 Canthodimorpha Davis, Scholtz & Harrison, 1999 Canthon Hoffmannsegg, 1817 Canthonella Chapin, 1930 Canthonidia Paulian, 1939 Canthonosoma MacLeay, 1871 Canthotrypes Paulian, 1939 Cantotius Bordat, 2022 Canudema Lacroix, 1994 Canuschiza Lacroix, 1999 Carinaulus Tesař, 1945 Carlschoenherria Bezděk, 2016 Carneiola Endrödi, 1974 Carneluttia Mikšić, 1976 Carneodon Özdikmen, 2009 Carneoryctes Özdikmen, 2009 Cartwrightia Islas, 1958 Castanochilus Ohaus, 1909 Castelnauphylla Lacroix, 1999 Catharsiocopris Balthasar, 1967 Catharsius Hope, 1837 Catoclastus Solier, 1851 Catrachia Coca-Abia & Robbins, 2006 Cavonus Sharp, 1875 Caymania Ratcliffe & Cave, 2010 † Ceafornotensis Woolley, 2016 Celidota Burmeister, 1842 Centrantyx Fairmaire, 1884 Centrochilus Krikken, 1976 Centrognathus Guérin-Méneville, 1840 Cephalocyclus Dellacasa & Gordon & Dellacasa, 1998 Cephalodesmius Westwood, 1842 Cephaloncheres Brenske, 1898 Cephaloschiza Moser, 1920 Ceramida Baraud, 1987 Ceraspis Le Peletier & Serville, 1828 Ceratochelus Dombrow, 2002 Ceratogonia Kolbe, 1899 Ceratolontha Arrow, 1948 Ceratophileurus Ohaus, 1911 Ceratoryctoderus Arrow, 1908 Cerhomalus Quedenfeldt, 1884 Ceroplophana Gestro, 1893 Cesamexico Koçak & Kemal, 2008 Cestradoretus Ohaus, 1912 Cetonia Fabricius, 1775 Chaetocosmetes Moser, 1917 Chaetodermina Heller, 1921 Chaetonyx Schaum, 1862 Chaetopisthes Westwood, 1847 Chaetopteroplia Medvedev, 1949 Chaetovalgus Moser, 1914 Chalcasthenes Arrow, 1937 Chalcentis Burmeister, 1844 Chalcochlamys Ohaus, 1905 Chalcocopris Burmeister, 1846 Chalcocrates Heller, 1903 Chalconotus Dejean, 1833 Chalcopharis Heller, 1902 Chalcoplethis Burmeister, 1844 Chalcosoma Hope, 1837 Chalcothea Burmeister, 1842 Chalcotheomima Mikšić, 1970 Chalepides Casey, 1915 Charadronota Burmeister, 1842 Chariochilus Brenske, 1898 Chariodactylus Moser, 1919 Chariodema Blanchard, 1850 Charioserica Brenske, 1899 Charitovalgus Kolbe, 1904 Chasmatopterus Dejean, 1821 Chasme Le Peletier & Serville, 1828 Chaunocolus Saylor, 1937 Cheilo Britton, 1990 Cheiragra MacLeay, 1864 Cheirodontus Britton, 1957 Cheirolasia Westwood, 1842 Cheironitis Lansberge, 1875 Cheiroplatys Hope, 1837 Cheirora Britton, 1957 Cheirotonus Hope, 1840 Cheleion Vardal & Forshage, 2010 Chelilabia Morón & Nogueira, 1998 Cherbezatina Lacroix, 1993 Cheronia Croissandeau, 1893 Chewia Legrand, 2004 Chilenopilus Smith & Mondaca, 2016 Chilodiplus Sharp, 1877 Chiloloba Burmeister, 1842 Chiloschiza Moser, 1924 Chilothorax Motschulsky, 1859 Chilotrogus Reitter, 1905 Chioneosoma Kraatz, 1891 Chipita Soula, 2008 Chiriquibia Bates, 1889 Chiron MacLeay, 1819 Chittius Tarasov, 2008 Chlaenobia Blanchard, 1850 Chloresthia Fairmaire, 1905 Chlorixanthe Bates, 1889 Chlorobapta Kraatz, 1880 Chlorocala Kirby, 1828 Chlorota Burmeister, 1844 Chnaunanthus Burmeister, 1844 Chondropyga Kraatz, 1880 Chondrorrhina Kraatz, 1880 Chromoptilia Westwood, 1842 Chromovalgus Kolbe, 1897 Chrysina Kirby, 1828 Chrysophora Le Peletier & Serville, 1828 Chrysoserica Brenske, 1897 Chthonobius Burmeister, 1847 Chuchina Soula, 2005 Cinacanthus Schmidt, 1914 Circellium Latreille, 1829 Clania Schein, 1956 Clastocnemis Burmeister, 1840 Clavipalpus Laporte, 1832 Cleptocaccobius Cambefort, 1984 Clerota Burmeister, 1842 Clilopocha Lea, 1914 Clinteria Burmeister, 1842 Clinterocera Motschulsky, 1857 Clinteroides Schoch, 1898 Clipadoretus Ohaus, 1941 Clithria Burmeister, 1842 Clitopa Erichson, 1847 Clomecyra Lacroix, 1999 Clypeasta Fairmaire, 1904 Clypeodrepanus Krikken, 2009 Clypeolontha Li & Yang, 1999 Clypeoserica Frey, 1970 Clyster Arrow, 1908 Cnemargulus Semenov, 1903 Cnematoplatys Schmidt, 1914 Cnemida Kirby, 1827 Cnemidophileurus Kolbe, 1910 Cnemisus Motschulsky, 1868 Cnemoschiza Decelle, 1979 Cocherellus Paulian, 1991 Cochinchidema Lacroix, 1996 Cochliotis Kolbe, 1894 Cochliotodes Burgeon, 1946 Coega Péringuey, 1902 Coelocorynus Kolbe, 1895 Coelocratus Burmeister & Schaum, 1841 Coelogenia Moser, 1913 Coelosis Hope, 1837 Coelotrachelus Schmidt, 1914 Coenochilus Schaum, 1841 Coenonycha Horn, 1876 Coenoryctoderus Prell, 1933 Coilodera Hope, 1831 Colacus Ohaus, 1910 Collagenus Ratcliffe & Hardy, 2005 Colobopteridius Clément, 1958 Colobopteroides Paulian, 1942 Colobopterus Mulsant, 1842 Colobostoma Blanchard, 1850 Colpochelyne Britton, 1987 Colpochila Erichson, 1843 Colpochilodes Blackburn, 1898 Colymbomorpha Blanchard, 1850 Comaserica Brenske, 1897 Comatapogonia Lacroix, 2008 Comencya Lacroix, 1993 Comophorina Strand, 1928 Comoramorphochelus Lacroix, 1997 Compsocephalus White, 1845 Compsodactylus Fuhrmann, 2012 Comythovalgus Kolbe, 1884 Conebius Fauvel, 1903 Coniopholis Erichson, 1847 Conioserica Brenske, 1900 Conradtia Kolbe, 1892 Copridaspidus Boucomont, 1920 Coprimorphus Mulsant, 1842 Copris Geoffroy, 1762 Coproecus Reiche, 1841 Coprophanaeus Olsoufieff, 1924 Coptochiroides Balthasar, 1938 Coptochirus Harold, 1859 Coptodactyla Burmeister, 1846 Coptognathus Burmeister, 1847 Coptomia Burmeister, 1842 Coptomiopsis Pouillaude, 1919 Coptorhina Hope, 1833 Corynophyllus Hope, 1845 Corynoserica Moser, 1918 Corynotrichius Kolbe, 1892 Corythoderus Klug, 1845 Coscinocephalus Prell, 1936 Cosmesthes Kraatz, 1880 Cosmiomorpha Saunders, 1852 Cosmiophaena Kraatz, 1898 Cosmovalgus Kolbe, 1897 Costelytra Given, 1952 Costinota Schürhoff, 1933 Cotalpa Burmeister, 1844 Cotinis Burmeister, 1842 Craniorphnus Kolbe, 1895 Craterocyphus Schmidt, 1914 Crathoplus Blanchard, 1851 Cremastocheilus Knoch, 1801 Crepischiza Brenske, 1903 † Cretaclopus Nikolajev, 2004 † Cretaegialia Nikolajev, 1993 † Cretomelolontha Nikolajev, 1998 † Cretonitis Nikolajev, 2007 † Cretorabaeus Nikolajev, 1995 † Cretoscarabaeus Nikolajev, 1995 † Cretoserica Nikolajev, 1998 Cryptaphodius Bordat, 2009 Cryptocanthon Balthasar, 1942 Cryptodontes Burmeister, 1847 Cryptodus MacLeay, 1819 Cryptoscatomaseter Gordon & Skelley, 2007 Cryptotrogus Kraatz, 1888 Cubidens Britton, 1992 Cunderdinia Lea, 1916 Cyclidiellus Krikken, 1976 Cyclidinus Westwood, 1874 Cyclidius MacLeay, 1838 Cyclocephala Dejean, 1821 Cyclomera Klug, 1855 Cylichnus Burmeister, 1844 Cylindrocrates Schein, 1958 Cymboptera Ohaus, 1923 Cymophorus Kirby, 1827 Cyobius Sharp, 1875 Cyphochilus Waterhouse, 1867 Cyphonistes Burmeister, 1847 Cyphonocephalus Westwood, 1842 Cyphonotus Fischer Von Waldheim, 1823 Cyphoserica Brenske, 1900 Cyptochirus Lesne, 1900 Cyriodera Burmeister, 1842 Cyriopertha Reitter, 1903 Cyrtocamenta Brenske, 1897 Cyrtolophus Fairmaire, 1895 Cyrtothyrea Kolbe, 1895 Cyrtotrochalus Brenske, 1900 Dactylopopillia Arrow, 1917 Daintreeola Howden & Storey, 2000 Dalatamala Prokofiev, 2013 Dalgopus Dechambre, 2001 Danielssonia Bordat & Howden, 1995 Dasygnathus MacLeay, 1819 Dasylepida Moser, 1913 Dasytrogus Reitter, 1902 Dasyus Le Peletier & Serville, 1828 Dasyvalgoides Endrödi, 1952 Dasyvalgus Kolbe, 1904 Debeckius Allsopp, 2022 Debutina Lacroix, 1993 Decellophylla Lacroix, 2005 Dechambrophylla Lacroix, 1999 Dedalopterus Sabatinelli & Pontuale, 1998 Degallieridium Vaz-de-Mello, 2008 Dellacasiellus Gordon & Skelley, 2007 Delopleurus Erichson, 1848 Delphinobius Fairmaire, 1900 Deltepilissus Pereira, 1949 Deltochilum Eschscholtz, 1822 Deltorhinum Harold, 1867 Demarziella Balthasar, 1961 Democrates Burmeister, 1847 Dendropaemon Perty, 1830 Denhezia Dechambre, 2006 Dentatoplia Lacroix, 1997 Denticnema Schein, 1959 Dentiheterochelus Dombrow, 2002 Dermolepida Arrow, 1941 Deroserica Moser, 1915 Descarpentriesia Ruter, 1964 Desertaclopus Ocampo & Mondaca, 2012 Desfontainesia Alexis & Delpont, 1999 Desicasta Thomson, 1878 Desmonyx Arrow, 1907 Diabroctis Gistel, 1857 Diaclaspus Brenske, 1896 Diacucephalus De Wailly, 1950 Dialithus Parry, 1849 Dialytellus Brown, 1929 Dialytes Harold, 1869 Dialytodius Gordon & Skelley, 2007 Diaphonia Newman, 1840 Diaphoroserica Péringuey, 1904 Diaphylla Erichson, 1847 Diaplochelus Burmeister, 1844 Diapterna Horn, 1887 Diastellopalpus Lansberge, 1886 Diastictus Mulsant, 1842 Dicaulocephalus Gestro, 1888 Dicellachilus Waterhouse, 1905 Dicentrines Burmeister, 1844 Dichecephala Brenske, 1895 Dichelhoplia Blanchard, 1850 Dichelomorpha Burmeister, 1855 Dichelonyx Harris, 1826 Dicheloschema Moser, 1924 Dichelus Le Peletier & Serville, 1828 Dicheros Gory & Percheron, 1833 Dichodontus Burmeister, 1847 Dichotomius Hope, 1838 Dichrosoma Kraatz, 1885 Dicrania Le Peletier & Serville, 1828 Dicranocara Frolov & Scholtz, 2003 Dicranocnemus Burmeister, 1844 Dicranoplia Reitter, 1903 Dicronocephalus Hope, 1831 Dicronorhina Hope, 1837 Didactylia Orbigny, 1896 Didrepanephorus Wood-mason, 1878 Digenethle Thomson, 1877 Digitonthophagus Balthasar, 1959 Dihymenonyx Gutiérrez, 1949 Dikellites Britton, 1990 Dilatatoplia Lacroix, 1998 Diloboderus Reiche, 1859 Dilochrosis Thomson, 1878 Dilophochila Bates, 1888 Dilortomaeus Bordat, 2009 Dinacoma Casey, 1889 Dinamoraza Lacroix, 1989 Dinarobina Lacroix, 1989 Dinoryctes Felsche, 1906 Diorygopyx Matthews, 1974 Dipelicus Hope, 1842 Diphrontis Gerstaecker, 1882 Diphucephala Le Peletier & Serville, 1828 Diphycerus Fairmaire, 1878 Diphydactylus Thomson, 1858 Diphyllomorpha Hope, 1843 Diploa Kolbe, 1892 Diplognatha Gory & Percheron, 1833 Diplotaxis Kirby, 1837 Diplotropis Boheman, 1857 Dirrhina Burmeister, 1842 Dischista Burmeister, 1842 Discopeltis Burmeister, 1842 Disphysema Harold, 1873 Djadjoua Lacroix, 1993 Djafouna Lacroix, 1993 Doaphius Balthasar, 1941 Doleroserica Péringuey, 1904 Dolerotarsa Péringuey, 1904 Dolichiomicroscelis Dombrow, 2003 Dolichoplia Lacroix, 1998 Dolichostethus Kolbe, 1892 Dorbignyolus Branco, 1989 Doryscelis Burmeister, 1842 Dorysthetus Blanchard, 1845 Doxocalia Brenske, 1897 Drepanocanthoides Schmidt, 1914 Drepanocanthus Péringuey, 1901 Drepanocerus Kirby, 1828 Drepanoplatynus Boucomont, 1921 Drogo Deschodt, Davis & Scholtz, 2016 Dudleyellus Bordat, 2009 Dungoorus Carne, 1958 Dwesasilvasedis Deschodt & Scholtz, 2008 Dymusia Burmeister, 1842 Dynastes MacLeay, 1819 Dyscinetus Harold, 1869 Dysphanochila Blackburn, 1898 Ebolowanius Dechambre, 1979 Eccoptomia Kraatz, 1880 Echyra Erichson, 1847 Ectinohoplia Redtenbacher, 1868 Ectinoplectron Ohaus, 1915 Eideria Neita & Ocampo, 2012 Elaphinis Burmeister, 1842 Elaphocera Gene, 1836 Elaphocerella Moser, 1919 Elcarmeniella Franz, 1955 Elpidus Péringuey, 1907 Emadiellus Schmidt, 1914 Embrithoplia Lacroix, 1998 Empecamenta Brenske, 1895 Empecta Erichson, 1847 Empectoides De Wailly, 1950 Emphania Erichson, 1847 Empycastes Gerstaecker, 1871 Enamillus Sharp, 1877 Enaria Erichson, 1847 Enarotadius Endrödi, 1971 Encya Dejean, 1833 Endebius Lansberge, 1880 Endoxazus Kolbe, 1892 Endroedianibe Chalumeau, 1981 Endroedyolus Scholtz & Howden, 1987 Enema Hope, 1837 Engertia Dalle Torre, 1912 Engistanoxia Lacroix, 2002 Engyopsina Britton, 1990 Ennearabdus Lansberge, 1874 Enoplotarsus Lucas, 1859 Enracius Dechambre, 1999 Enthora Erichson, 1847 Entypophana Moser, 1913 Entyposis Kolbe, 1894 Eocorythoderus Marumaya, 2012 Eodrepanus Barbero, Palestrini & Roggero, 2009 Eophileurus Arrow, 1908 Eosaulostomus Carne, 1956 Eotrichia Medvedev, 1951 Epactoides Olsoufieff, 1947 Epadoretus Reitter, 1903 Epectinaspis Blanchard, 1851 Epicaulis Dejean, 1833 Epichalcoplethis Bates, 1904 Epichrysus White, 1841 Epidrepanus Roggero, Barbero & Palestrini, 2015 Epilissus Reiche, 1841 Epipholis Moser, 1917 Epirinus Dejean, 1833 Epironastes Carne, 1957 Epistalagma Fairmaire, 1880 Epitrichius Tagawa, 1941 Epixanthis Burmeister, 1842 Erbmahcedius Hutchinson & Allsopp, 2021 Ercomoana Lacroix, 1994 Eremazus Mulsant, 1851 Eremobothynus Ohaus, 1910 Eremophygus Ohaus, 1910 Eremoplia Kolbe, 1914 Eremotrogus Kolbe, 1914 Eriesthis Burmeister, 1844 Erigavo Lacroix, 2006 Eriopeltastes Burmeister & Schaum, 1840 Erioscelis Burmeister, 1847 Eriphoserica Brenske, 1899 Eriulis Burmeister, 1842 Erlangeria Preiss, 1902 Erytodes Schmidt, 1917 Erytus Mulsant & Rey, 1870 Escarabaeus Zídek & Pokorný, 2011 Esymus Mulsant & Rey, 1869 Etiserica Péringuey, 1904 Eucamenta Péringuey, 1904 Euchilia Burmeister, 1842 Euchirus Burmeister & Schaum, 1840 Euchloropus Arrow, 1907 Euchroea Burmeister, 1842 Eucopidocaulus Prell, 1912 Eucranium Brullé, 1837 Eucyclophylla Waterhouse, 1875 Eudicella White, 1839 Eudinopus Burmeister, 1840 Eudolus Mulsant & Rey, 1869 Euetheola Bates, 1888 Euhemicyclium Bordat, 2003 Euheptaulacus Dellacasa, 1983 Euknemoplia Lacroix, 1997 Eulaiades Fairmaire, 1899 Eulepida Kolbe, 1894 Eulepidopsis Burgeon, 1946 Euligyrus Casey, 1915 Eumacronota Mikšić, 1976 Eunanus Ohaus, 1909 Euoniticellus Janssens, 1953 Euonthophagus Balthasar, 1959 Euorodalus Dellacasa, 1983 Euotophorus Dellacasa, Gordon & Dellacasa, 2016 Euparia Le Peletier & Serville, 1828 Euparixia Brown, 1927 Euparixoides Hinton, 1936 Euparotrix Stebnicka & Howden, 1996 Eupatorus Burmeister, 1847 Eupegylis Duvivier, 1892 Euphoresia Brenske, 1897 Euphoria Burmeister, 1842 Eupleurus Mulsant, 1842 Eupoecila Burmeister, 1842 Euranoxia Semenov, 1890 Euremina Westwood, 1867 Europtron Marseul, 1867 Euryaspis Blanchard, 1851 Eurychelus Blanchard, 1850 Euryomia Burmeister, 1842 Eurypeza Lacroix, 2006 Euryschiza Brenske, 1898 Eurysternus Dalman, 1824 Euryvalgus Moser, 1908 Eusaproecius Branco, 1989 Euselates Thomson, 1880 Euserica Reitter, 1896 Eutelesmus Waterhouse, 1880 Eutrichesis Waterhouse, 1882 Eutrichillum Martinez, 1967 Eutyctus Semenov, 1890 Evanos Castelnau, 1840 Evbrittonia Szito, 1993 Exanticheira Soula, 1998 Exaphodius Endrödi, 1960 Eximidema Keith & Reichenbach, 2018 Exochlorota Soula, 2002 Exochogenys Carne, 1958 Exolontha Reitter, 1902 Exomala Reitter, 1903 Exopholis Motschulsky, 1859 Exoptenomela Soula, 2002 Exostethus Britton, 1979 Exothyridium Soula, 2002 Exoxyomus Bordat, Mencl & Rakovič, 2012 Extenuoptyophis Smith & Mondaca, 2015 Faargia Martinez, 1982 Fairmairoplia Lacroix, 1997 Falsignambia Paulian, 1987 Falsotoclinius Keith & Montreuil, 2004 Faula Blanchard, 1850 Feeridium Vaz-de-Mello, 2008 Ferrerianus Dellacasa, Dellacasa & Gordon, 2007 Firminus Coca-Abia, 2003 Flaviellus Gordon & Skelley, 2007 Flechtmanniella Stebnicka, 1999 Fornasinius Bertoloni, 1852 Fossocarus Howden, 1961 Frankenbergerius Balthasar, 1938 Frenchella Blackburn, 1892 Fruhstorferia Kolbe, 1894 Fulvaphodius Bordat, 2022 Gametis Burmeister, 1842 Gametoides Antoine, 2005 Gamka Péringuey, 1904 Ganganomala Ratcliffe, Jameson & Zorn, 2018 Gariep Péringuey, 1907 Garreta Janssens, 1940 Gastromaladera Nomura, 1973 Gastroserica Brenske, 1897 Geniates Kirby, 1818 Geniatosoma Costa Lima, 1940 Genieridium Vaz-de-Mello, 2008 Genuchinus Westwood, 1874 Genuchus Kirby, 1825 Geomyphilus Gordon & Skelley, 2007 Geopsammodius Gordon & Pittino, 1992 Geotrogus Guérin-Méneville, 1842 Gibboryctes Endrödi, 1974 Giesbertiolus Howden, 1988 Gillaspytes Howden, 1980 Gilletellus Janssens, 1937 Gilletianus Balthasar, 1933 Glacilitarsus Matsumoto, 2014 Glaphyronyx Moser, 1924 Glaphyserica Brenske, 1899 Glenopopillia Lin, 1980 Globencya Lacroix, 1993 Globulischiza Frey, 1974 Glossocheilifer Blackburn, 1898 Glycosia Schoch, 1896 Glycyphana Burmeister, 1842 Glycyserica Brenske, 1899 Glyphoderus Westwood, 1838 Glyptoglossa Brenske, 1895 Glyptothea Bates, 1889 Glyptotheomima Mikšić, 1976 Gnaphalopoda Reiche, 1860 Gnaphalostetha Reiche, 1856 Gnathocera Kirby, 1825 Gnathogolofa Arrow, 1914 Gnatholabis Erichson, 1847 Gnorimella Casey, 1915 Gnorimidia Lansberge, 1887 Gnorimimelus Kraatz, 1880 Gnorimotrichius Krajčik, 2009 Gnorimus Le Peletier & Serville, 1828 Goiginus Endrödi, 1982 Goliathopsis Janson, 1881 Goliathus Lamarck, 1801 Golinca Thomson, 1878 Golofa Hope, 1837 Gonaphodiellus Schmidt, 1914 Gonaphodioides Dellacasa, Dellacasa & Gordon, 2012 Gonaphodiopsis Dellacasa, Dellacasa & Gordon, 2012 Gonaphodius Reitter, 1892 Gongrolophus Stebnicka & Howden, 1996 Goniaspidius Burmeister, 1844 Goniophileurus Kolbe, 1910 Goniorphnus Arrow, 1911 Goniorrhina Quedenfeldt, 1888 Gorditus Ratcliffe, 2010 Gordonius Skelley, Dellacasa & Dellacasa, 2009 Gouna Péringuey, 1902 Gracilaclopus Ocampo & Mondaca, 2012 Grammodoplia Lacroix, 1997 Grammopyga Kolbe, 1895 Grandaustralis Hutchinson & Moeseneder, 2013 Grandinaphodius Ziani, 2002 Granulopsammodius Rakovič, 1981 Graviengertia Prokofiev, 2016 Grebennikovius Mlambo, Scholtz & Deschodt, 2019 Griveaudella Lacroix, 1993 Gromphas Brullé, 1837 Gronocarus Schaeffer, 1927 Gryphonycha Péringuey, 1904 Guanyinaphodius Masumoto & Kiuchi, 2001 Guatemalica Neervoort Van De Poll, 1886 Gymnephoria Ratcliffe, 2019 Gymnetina Casey, 1915 Gymnetis MacLeay, 1819 Gymnogaster Blanchard, 1851 Gymnoloma Dejean, 1833 Gymnophana Arrow, 1910 Gymnopleurus Illiger, 1803 Gymnopyge Linell, 1896 Gymnoschiza Moser, 1914 Gynaecoserica Brenske, 1896 Gyronotus Lansberge, 1874 Gyroplia Brenske, 1893 Hadrodiplognatha Kraatz, 1898 Hadropechys Britton, 1990 Hadrops Britton, 1987 Hadrorhyparus Howden, 1995 Hadrosticta Kraatz, 1892 Halffterinetis Morón & Nogueira, 2007 Hamatoplectris Frey, 1967 Hammondantus Cambefort, 1978 Hamonthophagus Roggero, Dierkens, Barbero & Palestrini, 2017 Hansreia Halffter & Martinez, 1977 Haplidia Hope, 1837 Haplophileurus Kolbe, 1910 Haploscapanes Arrow, 1908 Haplosoma Semenov, 1890 Harmodactylus Péringuey, 1901 Harmogaster Harold, 1861 Haroldaphodius Bordat, 2003 Haroldiataenius Chalumeau, 1981 Haroldiellus Gordon & Skelley, 2007 Haroldius Boucomont, 1914 Harpechys Britton, 1957 Harpina Burmeister, 1844 Harposceles Burmeister, 1847 Hatamus Sharp, 1877 Hecistopsilus Kolbe, 1894 Hegemus Thomson, 1881 Helictopleurus Orbigny, 1915 Heliocopris Hope, 1837 Hellaserica Baraud & Nicolas, 1966 Hemiaspidius Krikken, 1982 Hemicamenta Brenske, 1897 Hemichaetoplia Baraud, 1986 Hemichalcothea Mikšić, 1970 Hemichnoodes Kraatz, 1880 Hemicorythoderus Tangelder & Krikken, 1982 Hemictenius Reitter, 1897 Hemicyclium Schmidt, 1914 Hemicyrthus Reiche, 1860 Hemiheterorrhina Mikšić, 1974 Hemilia Kraatz, 1880 Hemiphaedimus Mikšić, 1972 Hemipharis Burmeister, 1842 Hemiphileurus Kolbe, 1910 Hemiprotaetia Mikšić, 1963 Heptaulacus Mulsant, 1842 Heptelia Brenske, 1898 Heptophylla Motschulsky, 1857 Herculaisia Seilliere, 1910 Heteralleucosma Antoine, 1989 Heterochelus Burmeister, 1844 Heterochlorota Soula, 2002 Heteroclita Burmeister, 1842 Heteroclitopus Péringuey, 1901 Heterocnemis Albers, 1852 Heteroconus Kolbe, 1900 Heterocotinis Martinez, 1948 Heterocranus Bourgoin, 1919 Heterogeniates Ohaus, 1909 Heterogenius Moser, 1911 Heteroglobus Dupuis & Dechambre, 2008 Heterogomphus Burmeister, 1847 Heteroligus Kolbe, 1900 Heteronitis Gillet, 1911 Heteronychus Dejean, 1833 Heteronyx Guérin-Méneville, 1830 Heterophana Burmeister, 1842 Heterophthalmus Blanchard, 1851 Heteropseudinca Valck Lucassen, 1933 Heterorhina Westwood, 1842 Heteroschiza Moser, 1917 Heteroserica Brenske, 1899 Heterosoma Schaum, 1844 Heterosternus Dupont, 1832 Heterostriatus Dupuis, 2006 Heterotephraea Antoine, 2002 Heterotrochalus Moser, 1918 Heterovalgus Krikken, 1978 Hexataenius Fairmaire, 1891 Hexodon Olivier, 1789 Hiberasta Fairmaire, 1901 Hiekeianus Endrödi, 1978 Hieritis Burmeister, 1855 Hilyotrogus Fairmaire, 1886 Himalhoplia Sabatinelli, 1983 Hispanioryctes Howden & Endrodi, 1978 † Holcorobeus Nikritin, 1977 Holisonycha Péringuey, 1904 Holocanthon Martinez & Pereira, 1956 Holocephalus Hope, 1838 Holochelus Reitter, 1889 Holocnemus Brenske, 1894 Hologymnetis Martinez, 1949 Holomelia Brenske, 1891 Holopycnia Brenske, 1896 Holorhopaea Britton, 1978 Holotrichia Hope, 1837 Homalochilus Blanchard, 1851 Homaloserica Brenske, 1897 Homalotarsus Janssens, 1932 Homeochlorota Soula, 2006 Homocopris Burmeister, 1842 Homoeomorphus Burmeister, 1847 Homoeoschiza Kolbe, 1894 Homoiosternus Ohaus, 1901 Homolotropus MacLeay, 1871 Homonyx Guérin-Méneville, 1839 Homophileurus Kolbe, 1910 Homopliopsis Lacroix, 1998 Homothermon Ohaus, 1898 Homothyrea Kolbe, 1895 Hoplebaea Brenske, 1899 Hopleidos Künckel D'Herculais, 1887 Hoplia Illiger, 1803 Hopliopsis Blanchard, 1850 Hoplitovalgus Kolbe, 1904 Hoplochelus Blanchard, 1851 Hoplocnemis Harold, 1869 Hoplolontha Fairmaire, 1904 Hoplomaladera Nomura, 1974 Hoplopelidnota Bates, 1904 Hoplopus Laporte, 1832 Hoplopyga Thomson, 1880 Hoplopygothrix Schürhoff, 1933 Hoplorida Moser, 1918 Hoploryctoderus Prell, 1933 Hoplosternodes Burgeon, 1946 Hornietus Stebnicka, 2000 Hornosus Dellacasa, Dellacasa & Gordon, 2015 Horridocalia Endrödi, 1974 Hovachelus Fairmaire, 1897 Hovophileurus Arrow, 1911 Howdenocarus Hardy, 1978 Howdenypa Arnaud, 1993 Humblotania Lacroix, 1993 Hyalonthophagus Palestrini & Giacone, 1988 Hybaloides Quedenfeldt, 1884 Hybalus Dejean, 1833 Hybocamenta Brenske, 1898 Hyboschema Péringuey, 1901 Hyboserica Péringuey, 1904 † Hybosorites Nikolajev, 1996 Hybovalgus Kolbe, 1904 Hylamorpha Arrow, 1899 Hylobothynus Ohaus, 1910 Hymenochelus Reitter, 1890 Hymenoplia Eschscholz, 1830 Hypaspidius Arrow, 1899 Hyperius Fairmaire, 1878 Hyphelithia Kraatz, 1880 Hyphoryctes Blackburn, 1895 Hypocanthidium Balthasar, 1938 Hypolepida Britton, 1978 Hyposerica Brenske, 1897 Hypothyce Howden, 1968 Hypotrichia LeConte, 1861 Hypselogenia Burmeister, 1840 Iarupea Martinez, 1953 Iberoaphodius Rössner, 2018 Ichnestoma Gory & Percheron, 1833 Ictigaster Britton, 1986 Idaecamenta Péringuey, 1904 Idaeserica Péringuey, 1904 Idanastes Britton, 1987 Idiapogonia Arrow, 1916 Idiochelyna Arrow, 1916 Idionycha Arrow, 1932 Idiovalgus Arrow, 1910 Idutywa Péringuey, 1902 Ignambia Heller, 1916 Iguazua Stebnicka, 1997 Ihosyus Montreuil & Lacroix, 2020 Inanda Péringuey, 1902 Inca Le Peletier & Serville, 1828 Incala Thomson, 1857 Indieraligus Dechambre, 1979 Indoblackburneus Mencl, Král & Rakovič, 2019 Indognorimus Krikken, 2009 Indotrichius Krikken, 2009 Indovalgus Ricchiardi, 2012 Ingogius Endrödi, 1976 Ingrisma Fairmaire, 1893 Insimuloissacaris Smith & Mondaca, 2015 Iranotrogus Nikolajev, 2006 Irazua Ratcliffe, 2003 Iridisoma Delgado & Morón, 1991 Irrasinus Gordon & Skelley, 2007 Isacanthon Pacheco & Vaz-de-Mello, 2019 Ischiopsopha Gestro, 1874 Ischnopopillia Kraatz, 1892 Ischnoscelis Burmeister, 1842 Ischnostomiella Krikken, 1978 Ischnotarsia Kraatz, 1880 Ischnovalgus Kolbe, 1897 Isocopris Pereira & Martinez, 1960 Isonychus Mannerheim, 1828 Isoplia Burmeister, 1855 Issacaris Fairmaire, 1889 Ixodina Roth, 1851 Ixorida Thomson, 1880 Jalalabadia Balthasar, 1967 Jalisco Dellacasa, Gordon & Dellacasa, 2003 Jansonia Schürhoff, 1937 Janssensantus Paulian, 1976 Janssensellus Cambefort, 1975 Jossonthophagus Dierkens, Moretto & Génier, 2017 Joziratia Lacroix, 1993 Jumnos Saunders, 1839 Junkia Dalla Torre, 1913 † Juraclopus Nikolajev, 2005 Kabindeknomiosoma Lacroix, 2005 Kadlecia Frey, 1965 Kareiga Péringuey, 1902 Karolinella Minkina, 2017 Karooida Perissinotto, 2020 Kenyalepis Lacroix, 2011 Kenyatopadema Lacroix, 2009 Kerowagia Delpont, 1996 Kheper Janssens, 1940 Khoina Péringuey, 1902 Kibakoganea Nagai, 1984 Kirprellius Allsopp, 2022 Knysna Péringuey, 1902 Kolbeellus Jacobson, 1906 Komrina Lacroix, 1993 Koreoxyomus Kim, 1996 Korisaba Péringuey, 1902 Koshantschikovius Schmidt, 1914 Kraseophylla Péringuey, 1904 Krikkenius Branco, 1991 Kryzhanovskia Nikolajev & Kabakov, 1977 Kubousa Péringuey, 1902 Kuenckeliana Lacroix, 1989 Kurtops Roggero, Barbero & Palestrini, 2017 Labarrus Mulsant & Rey, 1869 Labroma Sharp, 1873 Laceratoplia Lacroix, 1997 Lachnodera Erichson, 1847 Lachnota Reitter, 1889 Lacroixidema Keith, 2000 Lacroixilepis Keith, 2007 Lagochile Hoffmannsegg, 1817 Lamellothyrea Krikken, 1980 Lamproserica Brenske, 1897 Lansbergia Ritsema, 1888 Laotrichia Keith, 2007 Larhodius Balthasar, 1963 Lasiexis Semenov-Tian-Shanskii & Medvedev, 1936 Lasiocala Blanchard, 1851 Lasiopsis Erichson, 1847 Lasioserica Brenske, 1897 Lasiotrichius Reitter, 1899 Lasiotropus Reitter, 1913 Latipalpus Moser, 1921 Latodrepanus Krikken, 2009 Lebbea Lacroix, 1993 Lecanoderus Kolbe, 1907 Lecanotrogus Kolbe, 1894 Lechorodius Gordon & Skelley, 2007 Leiopsammodius Rakovič, 1981 Lenosoma MacLeay, 1863 Leonotus Britton, 1986 Leoserica Fabrizi, Eberle & Ahrens, 2019 Leotrichillum Vaz-de-Mello, 2008 Lepanus Balthasar, 1966 Lepidiota Kirby, 1828 Lepidomela Kolbe, 1894 Lepidoserica Nikolajev, 1979 Lepidotrogus Kolbe, 1894 Lepischiza Moser, 1914 Lepisia Le Peletier & Serville, 1828 Lepithrix Le Peletier & Serville, 1828 Lepitrichula Schein, 1959 Lepivalgus Moser, 1914 Leptochristina Baraud & Branco, 1991 Leptocnemis Dombrow, 2001 Leptohoplia Saylor, 1935 Leptorhyparus Howden, 2003 Leribe Péringuey, 1904 Leucocelis Burmeister, 1842 Leucopholis Dejean, 1833 Leucophorus Brenske, 1892 Leucothyreus MacLeay, 1819 Leuretra Erichson, 1847 Liatongus Reitter, 1892 Lichnia Erichson, 1835 Lichniops Gutiérrez, 1946 Lichniopsoides Martinez, 1953 Licnostrategus Prell, 1933 Ligyrus Burmeister, 1847 Limarus Mulsant & Rey, 1869 Limepomera Lacroix, 1999 Linargius Rakovič & Mencl, 2012 Lindbergianus Petrovitz, 1961 Liogenys Guérin-Méneville, 1830 Lioglyptoxenus Pittino, 2006 Liomenochilus Hutchinson & Allsopp, 2022 Liostraca Burmeister, 1842 Liothorax Motschulsky, 1859 Liotrichius Kolbe, 1892 Liparetrus Guérin-Méneville, 1830 Lissadoretus Arrow, 1917 Lissogenius Schaum, 1844 Lissomelas Bates, 1889 Listrochelus Blanchard, 1851 † Lithanomala Nikolajev, 1992 † Lithoscarabaeus Nikolajev, 1992 Litocopris Waterhouse, 1891 † Lobateuchus Montreuil, Génier & Nel, 2010 Lobogeniates Ohaus, 1917 Loboparius Schmidt, 1914 Loboserica Kolbe, 1914 Lobovalgus Kolbe, 1897 Lomanoxia Martinez, 1951 Lomanoxoides Stebnicka, 1999 Lomaptera Gory & Percheron, 1833 Lonchothyrea Kolbe, 1895 Lonchotus Burmeister, 1847 Longaphodius Endrödi, 1980 Longicrura Frey, 1974 Lophodonitis Janssens, 1938 Lophorrhina Westwood, 1842 Lophorrhinides Perissinotto, Clennell & Beinhundner, 2019 Loraphodius Reitter, 1892 Loraspis Mulsant & Rey, 1869 Lorditomaeus Péringuey, 1901 Loricatarsa Frey, 1969 Lorkovitschia Mikšić, 1968 Lucassenia Olsoufieff, 1940 Luispenaia Martinez, 1972 Lunaphodius Balthasar, 1964 Lutera Westwood, 1875 Lutfius Özdikmen, 2009 Luxolinus Gordon & Skelley, 2007 Lycomedes Brême, 1844 Lyraphora Kraatz, 1880 Machala Lacroix, 1993 Macleayella Britton, 1990 Macraspis MacLeay, 1819 Macrocranius Schürhoff, 1935 Macrocyphonistes Ohaus, 1910 Macrodactylus Latreille, 1825 Macroderes Westwood, 1842 Macromina Westwood, 1874 Macronota Hoffmannsegg, 1817 Macronotops Krikken, 1977 Macroplia Brenske, 1898 Macropliopsis Lacroix, 1997 Macropoidelimus Morón, 1983 Macropoides Guérin-Méneville, 1844 Macropopillia Ohaus, 1905 Macroretroides Bordat, Dellacasa & Dellacasa, 2000 Macroretrus Péringuey, 1908 Macrotina Strand, 1934 Maculaphodius Gordon & Skelley, 2007 Madahoplia Lacroix, 1998 Madecorphnus Paulian, 1992 Madiana Ratcliffe & Romé, 2019 Madiniella Chalumeau & Gruner, 1976 Maechidinus Lea, 1919 Maechidius MacLeay, 1819 Maghrebotrogus Montreuil & Keith, 2022 Mahafalyanus Lacroix, 1993 Makalaka Péringuey, 1908 Makoanus Lacroix, 1993 Maladera Mulsant, 1871 Malagoniella Martinez, 1961 Malaia Heller, 1891 Malaisius Arrow, 1941 Malawinophylla Lacroix, 2013 Mallotarsus Blanchard, 1850 Manjarivolo Paulian, 1974 Manodactylus Moser, 1919 Manonychus Moser, 1919 Manopus Castelnau, 1840 Manoserica Moser, 1924 Marmarina Kirby, 1827 Marmylida Thomson, 1880 Marronus Coquerel, 1866 Martineziana Chalumeau & Özdikmen, 2006 Martinezidium Vaz-de-Mello, 2008 Mascarena Arrow, 1919 Masumotokoganea Hirasawa, 1992 Matthewsius Gunter & Weir, 2017 Mauromecistoplia Dombrow, 2002 Mausoleopsis Lansberge, 1882 Mawenzhena Alexis & Delpont, 2001 Mayataia Lacroix, 1993 Mazahuapertha Morón & Nogueira, 1998 Mecaspidiellus Antoine, 1997 Mecopelidnota Bates, 1904 Mecynodes Mulsant, 1870 Mecynorhina Hope, 1837 Medeotrogus Keith, 2001 Megaceras Hope, 1837 Megaceropsis Dechambre, 1976 Megacoryne Britton, 1987 Megalonitis Janssens, 1937 Megapertha (Reitter, 1903) Megaphonia Schürhoff, 1933 Megarhopaea Britton, 1978 Megasoma Kirby, 1825 Megasybacodes Kakizoe, Maruyama & Masumoto, 2019 Megateloides Landin, 1974 Megatelus Reitter, 1892 Megatharsis Waterhouse, 1891 Megathopa Eschscholtz, 1822 Megathoposoma Balthasar, 1939 Megistophylla Burmeister, 1855 Megistoplia Lacroix, 1997 Melanchroea Kraatz, 1900 Melanhyphus Fairmaire, 1881 Melanocamenta Brenske, 1899 Melanocanthon Halffter, 1958 Melanopopillia Lin, 1980 Melinopterus Mulsant, 1842 Mellissius Wollaston, 1869 Melolontha Fabricius, 1775 † Melolonthites Heer, 1847 Melolonthoides De Wailly, 1950 Mendidaphodius Reitter, 1901 Mendidius Harold, 1868 Menigionyx Arrow, 1941 Mentophilus Castelnau, 1840 Mericserica Brenske, 1897 Meridioclita Krikken, 1982 Merinanus Lacroix, 1993 Merogyrus Gordon & Skelley, 2007 Meroloba Thomson, 1880 Mesomerodon Ohaus, 1905 Mesontoplatys Motschulsky, 1863 Mesoserica Brenske, 1897 Mesosternus Morón, 1987 Mesotrochalus Kolbe, 1914 Messyrhus Rakovič & Král, 1997 † Mesydra Ponomarenko, 1977 Mesystoechus Waterhouse, 1878 Metacatharsius Paulian, 1939 Metadorodocia Machatschke, 1957 Metallesthes Kraatz, 1880 Metallopseudinca Valck Lucassen, 1933 Metanastes Arrow, 1911 Metapachylus Bates, 1889 Metaphileurus Kolbe, 1910 Metapogonia Lacroix, 2008 Metascelis Westwood, 1842 Metatrogus Britton, 1978 Metoporaphodius Frolov & Akhmetova, 2017 Meurguesia Ruter, 1969 Michaeloplia Lacroix, 1997 Micraegialia Brown, 1931 Micranomala Arrow, 1911 Micreuchilia Pouillaude, 1917 Microchalcothea Moser, 1910 Microchilus Blanchard, 1851 Microcoenus Britton, 1957 Microcopris Balthasar, 1958 Microdilochrosis Jákl, 2009 Microdoris Burmeister, 1844 Microlomaptera Kraatz, 1885 Microlontha Petrovitz, 1967 Microogenius Gutiérrez, 1951 Micropeltus Blanchard, 1842 Micropertha Baraud, 1991 Microphileurus Kolbe, 1910 Microphylla Kraatz, 1890 Microplidus Péringuey, 1902 Microplus Burmeister, 1844 Micropoecila Kraatz, 1880 Microrhopaea Lea, 1920 Microrutela Bates, 1904 Microryctes Arrow, 1908 Microserica Brenske, 1894 Microsericaria Nikolajev, 1979 Microtermitodius Pittino, 2006 Microteuchestes Landin, 1974 Microthopus Burmeister, 1855 Microtrochalus Brenske, 1900 Microvalgus Kraatz, 1883 Mictophileurus Ohaus, 1911 Milatrogus Montreuil & Keith, 2022 Milichus Péringuey, 1901 Mimadoretus Arrow, 1901 Mimaegialia Nikolajev, 2007 Mimela Kirby, 1823 Mimeoma Casey, 1915 Mimogeniates Martinez, 1964 Mimonthophagus Balthasar, 1963 Mimovalgus Arrow, 1944 Minidorysthetus Soula, 1998 Minisiderus Arrow, 1902 Minutoplia Lacroix, 1998 † Miolachnosterna Wickham, 1914 Miotemna Lacordaire, 1855 Mireia Ruter, 1953 Miridiba Reitter, 1902 Mitracamenta Brenske, 1903 Mitracephala Thomson, 1859 Mitrophorus Burmeister, 1844 Mnematidium Ritsema, 1888 Mnematium MacLeay, 1821 Modialis Fairmaire & Germain, 1860 Molynoptera Kraatz, 1897 Molynopteroides Antoine, 1989 † Mongolrobeus Nikolajev, 2004 Monochelus Le Peletier & Serville, 1828 Monoplistes Lansberge, 1874 Monotropus Erichson, 1847 Monteitheolus Howden & Storey, 2000 Monteithocanthon Gunter & Weir, 2017 Morettius Roggero, Dierkens, Barbero & Palestrini, 2017 Moriaphila Kraatz, 1880 Morokia Janson, 1905 Moroniella Ramírez-Ponce, 2015 Moronius Grossi & Vaz-de-Mello, 2015 Moronoserica Ahrens, Lukic & Liu, 2023 Moseriana Ruter, 1965 Mothon Semenov & Medvedev, 1927 Motuotrichius Huang & Chen, 2016 Mozartius Nomura & Nakane, 1951 Mucama Soula, 2002 Mucrencya Lacroix, 1993 Musurgus Vauloger, 1898 Mycernus Broun, 1904 Mycterophallus Neervoort Van De Poll, 1886 Myloxena Berg, 1881 Myloxenoides Martinez, 1975 Myodermides Ruter, 1964 Myodermidius Bourgoin, 1920 Myodermum Burmeister, 1840 Myrhessus Balthasar, 1955 Myrmecaphodius Martinez, 1952 Myrmecochilus Wasmann, 1900 Mysarus Petrovitz, 1962 Mystroceros Burmeister, 1842 Mzansica Perissinotto, 2020 Nagainokoganea Hirasawa, 1992 Namakwanus Scholtz & Howden, 1987 Namaphilus Deschodt & Davis, 2017 Nanaga Péringuey, 1902 Nanniscus Burmeister, 1844 Nannopopillia Kolbe, 1894 Nanorhopaea Britton, 1978 Nanos Westwood, 1842 Nanotermitodius Howden, 2003 Napoa Stebnicka, 1999 Narycius Dupont, 1835 Navigator Moeseneder & Hutchinson, 2016 Nayarita Morón & Nogueira, 1998 Neagolius Koshantschikov, 1912 Nebulasilvius Deschodt & Scholtz, 2008 Nedymoserica Brenske, 1900 Nefoncerus Saylor, 1938 Nematophylla Arrow, 1906 Nematoserica Arrow, 1917 Neocalaphodius Bordat, 1990 Neocamenta Burgeon, 1945 Neoclita Perissinotto, 2017 Neoclithria Neervoort Van De Poll, 1886 Neoclitopa Lacroix, 1997 Neocnecus Sharp, 1878 Neocolobopterus Landin, 1974 Neocorvicoana Ratcliffe & Mico, 2001 Neocorynophyllus Carne, 1985 Neodasygnathus Carne, 1957 Neodiapterna Dellacasa, 1984 Neoemadiellus Clément, 1986 Neogutierrezia Martinez, 1953 Neoheptaulacus Paulian & Villiers, 1941 Neoheteronyx Blackburn, 1890 Neohyphus Heller, 1896 Neomaladera Baraud, 1965 Neometanastes Yamaya, 2010 Neomyrhessus Minkina, 2023 Neomystroceros Alexis & Delpont, 1998 Neonastes Carne, 1957 Neonitis Péringuey, 1901 Neopegylis Lacroix, 2015 Neophaedimus Lucas, 1870 Neophaenognatha Allsopp, 1983 Neophyllotocus Blackburn, 1898 Neopsammodius Rakovič, 1986 Neorhyssemus Gordon & Pittino, 1992 Neorrhina Thomson, 1878 Neoryctes Arrow, 1908 Neosaproecius Branco, 1990 Neoscelis Schoch, 1897 Neoserica Brenske, 1894 Neosericania Miyake & Yamaya, 1994 Neotrichaphodioides Dellacasa, Dellacasa & Skelley, 2010 Neotrichiorhyssemus Rakovič & Král, 1997 Neotrichonotulus Dellacasa, Gordon & Dellacasa, 2004 Neovalgus Miyake, 1985 Nepaloserica Frey, 1965 Nephrodopus Sharp, 1873 Nepytis Erichson, 1842 Nesadmiralitia Prokofiev, 2016 Neso Blackburn, 1898 Nesohoplia Scott, 1912 Nesosisyphus Vinson, 1946 Nesovinsonia Martinez & Pereira, 1958 Nettelislasia Martinez, 1952 Neuquenodactylus Smith & Mondaca, 2015 Neuroserica Brenske, 1900 Nialaphodius Kolbe, 1908 Nialosternus Hollande & Therond, 1998 Nialus Mulsant & Rey, 1869 Nigroplia Dombrow, 2002 Nigrotrichia Matsumoto, 2016 Nimbacola Paulian, 1970 Nimbus Mulsant & Rey, 1869 Niphetophora Kraatz, 1883 Niphobleta Kraatz, 1880 Nipponaphodius Nakane, 1959 Nipponoagoliinus Ochi & Kawahara, 2001 Nipponoserica Nomura, 1973 Nipponovalgus Sawada, 1941 Nitiocellus Branco, 2010 Nitorellus Britton, 1957 Niuailan Allsopp & Hutchinson, 2020 Nobiellus Dellacasa & Dellacasa, 2005 Nobius Mulsant & Rey, 1869 Nolicus Petrovitz, 1962 Nosphisthis Blackburn, 1898 Nothophanus Heller, 1896 Notocaulus Quedenfeldt, 1884 Novapus Sharp, 1875 Nselenius (Holm & Perissinotto, 2004) Nuiba Prokofiev, 2016 Nunoidium Vaz-de-Mello, 2008 Nyassinus Westwood, 1879 Obaphodius Bordat, 1988 Oberthuroplia Lacroix, 1998 Ochicanthon Vaz-de-mello, 2003 Octocollis Moeseneder & Hutchinson, 2012 Octoplasia Brenske, 1892 Odochilus Harold, 1877 Odontacrossus Dellacasa, Král, Dellacasa & Bordat, 2014 Odontolochus Schmidt, 1917 Odontoloma Boheman, 1857 Odontolytes Koshantschikov, 1916 Odontoplia Fairmaire, 1897 Odontopsammodius Gordon & Pittino, 1992 Odontorrhina Burmeister, 1842 Odontotonyx MacLeay, 1871 Odontoxyomus Král, Rakovič & Mencl, 2015 Odontria White, 1846 Oedanomerus Waterhouse, 1875 Oedichira Burmeister, 1855 Oedipovalgus Kolbe, 1897 Oligophylla Kraatz, 1894 Omaloplia Schönherr, 1817 Omocnemus Schein, 1958 Omocrates Burmeister, 1844 Oncerus LeConte, 1856 Oncochirus Kolbe, 1891 Oniticellus Dejean, 1821 Onitis Fabricius, 1798 Onochaeta Erichson, 1847 Onoreidium Vaz-de-Mello, 2008 Onorius Frolov & Vaz-de-Mello, 2015 Ontherus Erichson, 1847 Onthobium Reiche, 1860 Onthophagus Latreille, 1802 Onychionyx Arrow, 1914 Onychoserica Moser, 1916 Onychosophrops Frey, 1972 Onychothecus Boucomont, 1912 Oocamenta Péringuey, 1904 Oogenius Solier, 1851 Ophropyx Britton, 1987 Oplognathus MacLeay, 1819 Oplostomus MacLeay, 1838 Opsitocus Britton, 1957 Orammoecius Král, Rakovič & Mencl, 2016 Oreoderus Burmeister, 1842 Oreogenoplia Lacroix, 1998 Oreotrogus Kolbe, 1897 Oreovalgus Kolbe, 1904 Orizabus Fairmaire, 1878 Orocanthus Endrödi, 1983 Oroclita Krikken, 1982 Orodaliscoides Schmidt, 1914 Orodaliscus Reitter, 1900 Oromus Mulsant & Rey, 1869 Orphnus MacLeay, 1819 Orrodoretus Ohaus, 1912 Orsilochus Burmeister, 1847 Orthocavonus Carne, 1957 Orthoserica Brenske, 1900 Orubesa Reitter, 1895 Oruscatus Bates, 1870 Oryctes Hellwig, 1798 † Oryctoantiquus Ratcliffe, Smith & Erwin, 2005 Oryctoderinus Endrödi, 1978 Oryctoderus Boisduval, 1835 Oryctomorphus Guérin-Méneville, 1830 Oryctophileurus Kolbe, 1910 Oscarinus Gordon & Skelley, 2007 Osmanius Branco & Baraud, 1988 Osmoderma Le Peletier & Serville, 1828 Otoclinius Brenske, 1896 Otophorus Mulsant, 1842 Ottokelleria D'Andretta & Martinez, 1957 Outenikwanus Scholtz & Howden, 1987 Outeniqua Péringuey, 1902 Ovomanonychus Costa, Cherman & Iannuzzi, 2020 Ovoserica Frey, 1968 Oxyadoretus Arrow, 1917 Oxyataenius Dellacasa & Stebnicka, 2001 Oxychirus Quedenfeldt, 1888 Oxycorythus Solsky, 1876 Oxyligyrus Arrow, 1908 Oxyomoides Dellacasa, Dellacasa & Gordon, 2016 Oxyomus Dejean, 1833 Oxypelta Pouillaude, 1920 Oxyserica Brenske, 1899 Oxysternon Castelnau, 1840 Oxythyrea Mulsant, 1842 Ozodius Skelley, 2007 Pachacama Soula, 2006 Pachnessa Brenske, 1894 Pachnoda Burmeister, 1842 Pachnodoides Alexis & Delpont, 2002 Pachrodema Blanchard, 1851 Pachycamenta Brenske, 1895 Pachychilecamenta Brenske, 1903 Pachycnema Le Peletier & Serville, 1828 Pachycolus Erichson, 1847 Pachydema Laporte, 1832 Pachydemocera Reitter, 1902 Pachyderoserica Moser, 1920 Pachygastra Germar, 1848 Pachylomera Griffith & Pidgeon, 1831 Pachyoryctes Arrow, 1908 Pachypoides Fairmaire, 1884 Pachypus Dejean, 1821 Pachyrhinadoretus Ohaus, 1912 Pachyserica Brenske, 1897 Pachysoma MacLeay, 1821 Pachystethus Blanchard, 1851 Pachytephraea De Palma & Malec, 2020 Pachytricha Hope, 1841 Pacuvia Curtis, 1845 Palacephala Lacroix, 1999 Palaeophileurus Kolbe, 1910 Paleira Reiche, 1871 Paleopragma Thomson, 1880 † Paleotrichius Poinar, 2011 Palnia Stebnicka, 1985 Panelus Lewis, 1895 Panglaphyra Kraatz, 1880 Panotrogus Reitter, 1902 Pantodinus Burmeister, 1847 Pantolia Burmeister, 1842 Papuana Arrow, 1911 Parabodilus Hollande & Thérond, 1998 Parabyrsopolis Ohaus, 1915 Paracamenta Péringuey, 1904 Paracanthon Balthasar, 1938 Paraceratochelus Dombrow, 2002 Parachilia Burmeister, 1842 Parachlorota Soula, 2002 Parachorius Harold, 1873 Parachrysina Bates, 1888 Paraclitopa Waterhouse, 1875 Paracoptochirus Balthasar, 1963 Paracorythoderus Wasmann, 1918 Paracotalpa Ohaus, 1915 Paracrossidius Balthasar, 1932 Paracryptocanthon Howden & Cook, 2002 Paracyclidius Howden, 1971 Paradeloparius Landin, 1959 Paradidactylia Balthasar, 1937 Paradoaphius Bordat, 2022 Paradorodocia Machatschke, 1957 Paradorysthetus Soula, 1998 Paraegidium Vulcano, Pereira & Martinez, 1966 Paragnorimus Becker, 1910 Paragymnopleurus Shipp, 1897 Paraheteronyx Moser, 1924 Paraheterosternus Morón, 1983 Paraholotrichia Lacroix & Coache, 2017 Paraixodina Roggero, Barbero & Palestrini, 2015 Paralleucosma Antoine, 1989 Parallidiostoma Ocampo & Colby, 2009 Paramacraspis Ohaus, 1915 Paramaladera Nomura, 1974 Paramicroplus Lacroix, 1998 Parammoecius Seidlitz, 1888 Paramorphochelus Lacroix, 1997 Paranelaphinis Antoine, 1988 Paranimbus Schmidt, 1914 Paranodon Cockerell, 1905 Paranomala Casey, 1915 Parapetiia Martinez, 1961 Paraphacosomoides Balthasar, 1969 Paraphileurus Endrödi, 1978 Paraphytus Harold, 1877 Parapilinurgus Arrow, 1910 Paraplectrone Mikšić, 1985 Paraplesiataenius Chalumeau, 1992 Parapoecilophila Hauser, 1904 Paraprotaetia Moser, 1907 Parapsammodius Verdu, Stebnicka & Galante, 2006 Paraptenomela Soula, 2002 Parapucaya Prell, 1934 Pararhabdotis Kraatz, 1899 Pararhopaea Blackburn, 1911 Pararhynchocephala Paulian, 1991 Pararhyssemus Balthasar, 1955 Parascatonomus Paulian, 1932 Paraschizognathus Ohaus, 1904 Parasciton Britton, 1990 Paraserica Reitter, 1896 Parastasia Westwood, 1842 Parastraella Antoine, 2005 Parasymmela Pacheco, Monné, Vaz-de-Mello & Ahrens, 2022 Parataenius Balthasar, 1961 Paratelaugis Ohaus, 1915 Parateuchus Shipp, 1895 Parathyridium Ohaus, 1915 Paratrichaphodius Bordat, 2013 Paratrichius Janson, 1881 Paratriodonta Baraud, 1962 Parautoserica Lacroix, Coache & Filippi, 2022 Paraxeloma Holm, 1988 Pardalosus Gordon & Skelley, 2007 Parelaphinis Marais & Holm, 1989 Parepixanthis Kraatz, 1893 Pareriesthis Moser, 1918 Pareteronychus Prell, 1934 Pareuchilia Kraatz, 1880 Parhomonyx Ohaus, 1915 Parhoplognathus Ohaus, 1915 Parisolea Bates, 1888 Parisomorphus Schaufuss, 1890 Paroniticellus Balthasar, 1963 Paronthobium Paulian, 1984 Paronyx Britton, 1990 Paroplia Fairmaire, 1903 Paroryctoderus Dechambre, 1994 Parthenoserica Brenske, 1899 Parvuhowdenius Deschodt & Scholtz, 2008 Pasaphylla Péringuey, 1904 Passaliolla Balthasar, 1945 Patatra Soula, 2008 Paulianellus Balthasar, 1938 Paulosawaya Martinez & D'andretta, 1956 Peckolus Scholtz & Howden, 1987 Pectinichelus Ballion, 1871 Pectinosoma Arrow, 1913 Pedaria Laporte, 1832 Pedaridium Harold, 1868 Pedinorrhina Kraatz, 1880 Pedinotrichia Matsumoto, 2016 Pegylis Erichson, 1847 Pelidnota MacLeay, 1819 † Pelidnotites Cockerell, 1920 Peltonotus Burmeister, 1847 Penalus Paulian, 1985 Pentacoryna Moser, 1926 Pentanomala Ohaus, 1919 Pentaphylla De Wailly, 1950 Pentecamenta Brenske, 1896 Pentelia Brenske, 1891 Penthima Kraatz, 1892 Pentodina Endrödi, 1968 Pentodon Hope, 1837 Pentodontoschema Péringuey, 1901 Peotoxus Krikken, 1983 Peperonota Westwood, 1847 Pereiraidium Vaz-de-Mello, 2008 Pericamenta Péringuey, 1904 Periclitopa Brenske, 1896 Pericoptus Burmeister, 1847 Periphanesthes Kraatz, 1880 Periproctoides Lacroix, 2006 Periproctus Kolbe, 1910 Periserica Brenske, 1898 Perissosoma Waterhouse, 1875 Peritrichia Burmeister, 1844 Perrindema Lacroix, 1997 Peruquime Mondaca & Valencia, 2016 Petinopus Blackburn, 1898 Petrovitzia Mikšić, 1965 Petrovitzius Rakovič, 1979 Phaeadoretus Reitter, 1903 Phaeaphodius Reitter, 1892 Phaedimus Waterhouse, 1841 Phaedotrogus Paulian, 1985 Phaenognatha Hope, 1841 Phaenomeris Hope, 1833 Phalacronothus Motschulsky, 1859 Phalangogonia Burmeister, 1844 Phalangonyx Reitter, 1889 Phalangosoma Quedenfeldt, 1884 Phalops Erichson, 1848 Phanaeus MacLeay, 1819 Phaneresthes Kraatz, 1894 Pharaonus Blanchard, 1851 Pharaphodius Reitter, 1892 Phiara Brenske, 1897 Philacelota Heller, 1900 Phileucourtus Dechambre, 2008 Phileurus Latreille, 1807 Philgertia Zídek, 2021 Philistina MacLeay, 1838 Philochloenia Dejean, 1833 Philoscaptus Brèthes, 1919 Philoserica Brenske, 1897 Phobetus LeConte, 1856 Pholeoaphodius Bordat, 2009 Pholidochris Kolbe, 1894 Phonopleurus Moser, 1919 Phonotaenia Kraatz, 1880 Phorine Britton, 1987 Photyna Brenske, 1897 Phoxomela Schaum, 1844 Phoxomeloides Schoch, 1898 Phycochus Broun, 1886 Phylliocephala Blackburn, 1890 Phyllocamenta Moser, 1918 Phyllochlaenia Blanchard, 1846 Phyllococerus Waterhouse, 1876 Phyllognathus Eschscholtz, 1830 Phyllopertha Stephens, 1830 Phyllophaga Harris, 1826 Phylloserica Brenske, 1899 Phyllotocidium Blackburn, 1898 Phyllotocus Fischer Von Waldheim, 1823 Phyllotrochalus Brenske, 1900 Phytholaema Blanchard, 1851 Pichica Soula, 2002 Pileotrichius Bourgoin, 1921 Pilinopyga Kraatz, 1888 Pilinurgus Burmeister, 1842 Pimelomera Moser, 1924 Pimelopus Erichson, 1842 Pinacopodius Branco, 1991 Pinacotarsus Harold, 1875 Piscoperus Ratcliffe & Giraldo, 2014 Pittinius Rakovič & Král, 1997 Placodidus Péringuey, 1900 Plaesioserica Brenske, 1899 Plagiogonus Mulsant, 1842 Planolinellus Dellacasa & Dellacasa, 2005 Planolinoides Dellacasa & Dellacasa, 2005 Planolinus Mulsant & Rey, 1869 Planophileurus Chapin, 1932 Platychelus Burmeister, 1844 Platycoelia Dejean, 1833 Platyderides Schmidt, 1917 Platygenia MacLeay, 1819 Platygeniops Krikken, 1978 Platynocephalus Westwood, 1854 Platyonitis Janssens, 1942 Platyphileurus Ohaus, 1910 Platyrutela Bates, 1888 Platysodes Westwood, 1874 Platytomus Mulsant, 1842 Plectris Le Peletier & Serville, 1828 Plectrodes Horn, 1867 Plectrone Wallace, 1867 Pleistophylla Péringuey, 1904 Pleophylla Erichson, 1847 Plesiocamenta Lacroix & Montreuil, 2019 Plesiopalacephala Lacroix, 2006 Plesiorutela Jameson, 1997 Plesiosternus Morón, 1983 Pleuraphodius Schmidt, 1914 Pleuronitis Lansberge, 1875 Pleuronota Kraatz, 1892 Pleurophorus Mulsant, 1842 Plochilia Fairmaire, 1896 Plochiliana Ruter, 1978 Plotopuserica Brenske, 1899 Plusioserica Brenske, 1899 Podalgus Burmeister, 1847 Podischnus Burmeister, 1847 Podolasia Harold, 1869 Podopholis Moser, 1915 Podopogonus Moser, 1917 Podostena Howden, 1997 Podotenoides Paulian, 1942 Podotenus Schmidt, 1914 Podovalgus Arrow, 1910 Poecilopharis Kraatz, 1880 Pogoniotarsus Kraatz, 1880 Pogonopus Arrow, 1910 Pogonotarsus Burmeister, 1842 Pollaplonyx Waterhouse, 1875 Polyenaria De Wailly, 1950 Polyphylla Harris, 1841 Polyphyllum Blanchard, 1851 Polyplastus Janson, 1888 Polystalactica Kraatz, 1882 Popillia Dejean, 1821 Porphyronota Burmeister, 1842 Prionoryctes Arrow, 1911 Priscorrhina Krikken, 1984 † Priscyparus Skelley, 2021 Priska Jákl, 2018 Pristerophora Harold, 1869 Proagoderus Lansberge, 1883 Proagopertha Reitter, 1903 Proagosternus Blanchard, 1851 Problerhinus Deyrolle, 1864 Proborhinus Britton, 1988 Prochelyna Erichson, 1847 Proctophanes Harold, 1861 Proculigyrus López-García & Deloya, 2022 Prodontria Broun, 1904 Prodoretus Brenske, 1893 Promacropoides Sigwalt, 1987 † Prophaenognatha Bai, Ren & Yang, 2011 Propomacrus Newman, 1836 Prosphileurus Kolbe, 1905 Protaetia Burmeister, 1842 Protaetiomorpha Mikšić, 1968 Protelura Britton, 1987 Proteroschiza Brenske, 1896 Protochilus Krikken, 1976 Protoclita Krikken, 1978 Protoclitopa Decelle, 1979 † Protopalnia Nikolajev, 2007 † Prototrox Nikolajev, 2000 † Psammaegialia Nikolajev, Wang & Zhang, 2014 Psammodaphodius Endrödi, 1976 Psammodius Fallen, 1807 Psammoporus Thomson, 1859 Psammorpha Stebnicka, 1994 Psednoserica Brenske, 1899 Pseudachloa Péringuey, 1904 Pseudacrossus Reitter, 1892 Pseudadoretus Semenov, 1890 Pseudagenius Heller, 1923 Pseudagolius Schmidt, 1914 Pseudalleucosma Antoine, 1989 Pseudastoxenus Bourgoin, 1921 Pseudataenius Brown, 1927 Pseudenaria Fairmaire, 1901 Pseudencya Lacroix, 1991 Pseudepixanthis Kraatz, 1880 Pseuderytus Hollande & Thérond, 1998 Pseudesymus Orbigny, 1896 Pseudheterochelus Schein, 1959 Pseudholophylla Blackburn, 1911 Pseudignambia Paulian, 1984 Pseudinca Kraatz, 1880 Pseudisonychus Frey, 1971 Pseudoagoliinus Bordat, 1994 Pseudoahermodontus Dellacasa & Dellacasa, 1997 Pseudoanticheiroides Soula, 1998 Pseudoblackburneus Bordat, 2016 Pseudoblitopertha Keith, 2000 Pseudocanthon Bates, 1887 Pseudocavonus Blackburn, 1890 Pseudochalcothea Ritsema, 1882 Pseudochalcotheomima Mikšić, 1985 Pseudochironitis Ferreira, 1977 Pseudochlorota Ohaus, 1905 Pseudoclinteria Kraatz, 1882 Pseudoclithria Neervoort Van De Poll, 1886 Pseudocoelotrachelus Dellacasa, Dellacasa & Gordon, 2013 Pseudocongella Moser, 1924 Pseudocopris Ferreira, 1960 Pseudocotalpa Hardy, 1971 Pseudocyrtolophus Frolov & Montreuil, 2019 Pseudodiceros Mikšić, 1974 Pseudodicrania Gutiérrez, 1950 Pseudodiplotaxis Nonfried, 1894 Pseudodorysthetus Soula, 1998 Pseudodrepanocanthus (Bordat, 1993) Pseudogeniates Ohaus, 1910 Pseudogonaphodiellus Dellacasa, Gordon & Dellacasa, 2007 Pseudoheptaulacus Dellacasa & Dellacasa, 2000 Pseudoheteronyx Blackburn, 1892 Pseudohomonyx Arrow, 1908 Pseudohypaspidius Soula, 1998 Pseudoliogenys Moser, 1919 Pseudomacraspis Ohaus, 1903 Pseudomaladera Nomura, 1974 Pseudomesystoechus Ohaus, 1912 Pseudomicroplus Lacroix, 1998 Pseudomothon Pittino, 1984 Pseudomyrhessus Minkina, 2023 Pseudonthobium Paulian, 1984 Pseudooxythyrea Baraud, 1985 Pseudopachydema Balthasar, 1930 Pseudopanotrogus Petrovitz, 1969 Pseudoparoplia Lacroix, 1997 Pseudopectinosoma Katovich, 2011 Pseudopedaria Felsche, 1904 Pseudoperiproctus Keith & Reichenbach, 2018 Pseudopharaphodius Bordat, 1990 Pseudopholis Duvivier, 1892 Pseudopilinurgus Moser, 1918 Pseudopodotenus Dellacasa, 1990 Pseudoptenomela Soula, 2002 Pseudorphnus Benderitter, 1913 Pseudoryctes Sharp, 1873 Pseudosaproecius Balthasar, 1941 Pseudoscaptobius Krikken, 1976 Pseudoschizognathus Ohaus, 1904 Pseudoschizonycha Keith, 2006 Pseudoserica Guérin-Méneville, 1830 Pseudosericania Kobayashi, 1980 Pseudostegopterus Ricchiardi, 2015 Pseudostereomera Bordat & Howden, 1995 Pseudosymmachia Dalle Torre, 1912 Pseudosyrichthus Péringuey, 1901 Pseudotephraea Kraatz, 1882 Pseudotetraclipeoides Dellacasa, Dellacasa & Gordon, 2015 Pseudoteuchestes Dellacasa, 1986 Pseudothyridium Soula, 2002 Pseudotocama Keith, 2008 Pseudotorynorrhina Mikšić, 1967 Pseudotrematodes Jacquelin Du Val, 1859 Pseudotrigonocnemis Keith, 2007 Pseudotrochalus Quedenfeldt, 1884 Pseudouteniqua Schein, 1959 Pseudoxyomus Petrovitz, 1962 Pseustophylla Péringuey, 1904 Psilocnemis Burmeister, 1842 Psilodontria Broun, 1895 Psilonychus Burmeister, 1855 Psilopholis Brenske, 1892 Ptenomela Bates, 1888 Pterobius Murayama, 2010 Ptychodesthes Kraatz, 1883 Ptyophis Redtenbacher, 1868 Pubinus Mulsant & Rey, 1869 Pucaya Ohaus, 1910 Puelchesia Ocampo & Smith, 2006 Pukupuku Muramoto, 2006 Pusiodactylus Smith, 2008 Pycnopanelus Arrow, 1931 Pycnoschema Thomson, 1859 Pygora Burmeister, 1842 Pygovalgus Kolbe, 1884 Pyronota Boisduval, 1835 Pyrrhopoda Kraatz, 1880 Qingaphodius Král, 1997 Rabula Péringuey, 1902 Raceloma Thomson, 1877 Rakovicius Pittino, 2006 Ramilia Kolbe, 1894 Ravautiana Lacroix, 1991 Raysymmela Saylor, 1947 Rectoscutaria Schein, 1958 Redotus Fairmaire, 1860 Reineria Mikšić, 1968 Renaudiana Lacroix, 1993 Renaudius Balthasar, 1951 Renorphnus Frolov & Montreuil, 2009 Repsimus MacLeay, 1819 Rhabdopholis Burmeister, 1855 Rhabdotis Burmeister, 1842 Rhabdotops Krikken, 1981 Rhachistoplia Lacroix, 1997 Rhadinolontha Arrow, 1930 Rhadinotaenia Kraatz, 1900 Rhagopteryx Burmeister, 1842 Rhamphadoretus Ohaus, 1912 Rhamphorrhina Klug, 1855 Rhinarion Ruter, 1965 Rhinaspis Perty, 1833 Rhinocerotopsis Maruyama, 2009 Rhinocoeta Burmeister, 1842 Rhinyptia Burmeister, 1844 Rhizogeniates Ohaus, 1909 Rhizoplatodes Péringuey, 1901 Rhizoplatys Westwood, 1841 Rhizoproctus Kolbe, 1894 Rhizotrogus Latreille, 1825 Rhodaphodius Adam, 1994 Rhomborhina Hope, 1837 Rhopaea Erichson, 1847 Rhynchadoretus Ohaus, 1913 Rhynchapogonia Lacroix, 2008 Rhynchocephala Fairmaire, 1883 Rhynchoserica Burgeon, 1942 Rhyparus Agassiz, 1846 Rhysothorax Bedel, 1911 Rhyssemodes Reitter, 1892 Rhyssemorphus Clouët, 1900 Rhyssemus Mulsant, 1842 Rhyxicephalus Moser, 1921 Rhyxiderma Moser, 1924 Rhyxiphloea Burmeister, 1842 Rigoutorum Hutchinson & Moeseneder, 2022 Rimuloplia Lacroix, 1997 Robinsonelliana Lacroix, 1996 Rubilepis De Wailly, 1950 Rufotrichia Matsumoto, 2016 Rugaphodius Gordon & Skelley, 2007 Rugopertha Machatschke, 1957 Rutela Latreille, 1802 Rutelarcha Waterhouse, 1874 Rutelisca Bates, 1888 Ruteloryctes Arrow, 1908 Ruteraetia Krikken, 1980 Sabatinellius Montreuil & Lacroix, 2020 Sabatinelloplia Lacroix, 1998 Saccharoscaptus Morón & Grossi, 2015 Sahlbergianus Dellacasa & Dellacasa, 2000 Saphobiamorpha Brookes, 1944 Saphobius Sharp, 1873 Saprolochus Stebnicka & Galante, 2007 Saprositellus Balthasar, 1967 Saprosites Redtenbacher, 1857 Saprovisca Stebnicka, 1993 Saprus Blackburn, 1904 Sariangus Rakovič & Mencl, 2011 Sarophorus Erichson, 1847 Sarothromerus Blackburn, 1907 Saulostomus Waterhouse, 1878 Sauvagesinella Paulian, 1934 Sayloria Frey, 1975 Scabrostomus Gordon & Skelley, 2007 Scapanes Burmeister, 1847 Scapanoclypeus Evans, 1987 Scaphorhina Quedenfeldt, 1884 Scaphorhinadoretus Ohaus, 1912 Scaptobius Schaum, 1841 Scaptocnemis Péringuey, 1901 Scarabaeolus Balthasar, 1965 Scarabaeus Linnaeus, 1758 Scatimus Erichson, 1847 Scatonomus Erichson, 1835 Scatrichus Génier & Kohlmann, 2003 Sceliages Westwood, 1837 Scelidothrix Dombrow, 2001 Scelophysa Burmeister, 1844 Schaefferellus Gordon & Skelley, 2007 Scheinia Ruter, 1957 Schismatocera Gautier Des Cottes, 1872 Schizadoretus Arrow, 1917 Schizochelus Blanchard, 1850 Schizognathus Fischer Von Waldheim, 1823 Schizonycha Dejean, 1833 Schizorhina Kirby, 1825 Schochidia Berg, 1898 Scitala Erichson, 1842 Scitaloides Britton, 1990 Sciton Blackburn, 1892 Scybalocanthon Martinez, 1948 Scybalophagus Martinez, 1953 Scythrodes Broun, 1886 Scythropesthes Kraatz, 1880 Sebakwe Péringuey, 1904 Sebaris Castelnau, 1840 Selaserica Brenske, 1897 Selomothus Fairmaire, 1891 Selviria Stebnicka, 1999 Semanopterus Hope, 1847 Semienaria De Wailly, 1950 Serica MacLeay, 1819 Sericania Motschulsky, 1860 Sericesthis Boisduval, 1835 Sericoides Guérin-Méneville, 1839 Sericospilus Sharp, 1882 Serraphodius Kabakov, 1996 Setiserica Miyake, 2002 Setodius Gordon & Skelley, 2007 Setylaides Stebnicka, 1994 Siamaphodius Masumoto, 1991 Siamophylla Keith, 2005 Sicardia Reitter, 1896 Sigorus Mulsant & Rey, 1869 Silluvia Landin, 1949 Silvaphilus Roets & Oberlander, 2010 Silvinha Vaz-de-Mello, 2008 Simogonius Harold, 1871 Simorrhina Kraatz, 1886 Sinaphodius Červenka, 1994 Sinapisoma Boucomont, 1928 Singhala Blanchard, 1851 Sinodiapterna Dellacasa, 1986 Sinodrepanus Simonis, 1985 Siralus Dechambre, 1998 Sisyphus Latreille, 1807 Sitiphus Fairmaire, 1894 Skelleyanus Dellacasa, Dellacasa, Gordon & Stebnicka, 2011 Smicorhina Westwood, 1847 Socotraproctus Král, Sehnal & Bezděk, 2012 Somalibia Lansberge, 1882 Somatoserica Brenske, 1899 Sophrops Fairmaire, 1887 Sorocha Soula, 2006 Spaniolepis Kolbe, 1894 Sparrmannia Castelnau, 1840 Spathoschiza Arrow, 1902 Spelaiorrhina Lansberge, 1886 Sphaeraphodius Kakizoe, Jiang & Wang, 2021 Sphaerorutela Jameson, 1997 Sphaeroscelis Burmeister, 1855 Sphaerotrochalus Brenske, 1900 Sphecoserica Brenske, 1899 Sphinctovalgus Kolbe, 1904 Sphodroxia Kraatz, 1890 Sphyrocallus Sharp, 1877 Spilophorus Westwood, 1848 Spilopopillia Kraatz, 1892 Spinanomala Ohaus, 1910 Spinochlamys Machatschke, 1970 Spodistes Burmeister, 1847 Spodochlamys Burmeister, 1855 Stalagmosoma Burmeister, 1842 Stebnickiella Skelley, 2007 Stegopterus Burmeister & Schaum, 1840 Stenochelyne Britton, 1990 Stenocrates Burmeister, 1847 Stenonota Fairmaire, 1889 Stenopegylis Arrow, 1943 Stenopisthes Moser, 1913 Stenoserica Brenske, 1900 Stenosophrops Nomura, 1977 Stenosternus Karsch, 1881 Stenotarsia Burmeister, 1842 Stenotothorax Schmidt, 1914 Stephanopholis Brenske, 1896 Stephanorrhina Burmeister, 1842 Stereomera Arrow, 1905 Stethaspis Hope, 1837 Stethodesma Bainbridge, 1840 Stethopseudinca Valck Lucassen, 1933 Stigmatoplia Dombrow, 2001 Stilbotrochalus Kolbe, 1914 Stiptocnemis Branco, 1989 Stiptopodius Harold, 1871 Stiptotarsus Branco, 1989 Stomanomala Kolbe, 1897 Storeyus Hasenpusch & Moeseneder, 2010 Straliga Fairmaire, 1901 Strategus Hope, 1837 Streblopus Lansberge, 1874 Strigoderma Burmeister, 1844 Strigodius Gordon & Skelley, 2007 Stripsipher Gory & Percheron, 1833 Strumadoretus Ohaus, 1912 Subrinus Mulsant & Rey, 1869 Sugrames Reitter, 1894 Sukelus Branco, 1992 Sulcophanaeus Olsoufieff, 1924 Suntemnonycha Péringuey, 1904 Supertermitoderus Mencl, 2011 Surutu Martinez, 1955 Sussorca Paulian, 1954 Sybacodes Fairmaire, 1896 Sybax Boheman, 1857 Sylvicanthon Halffter & Martinez, 1977 Symmela Erichson, 1835 Symphodon Schmidt, 1914 Synacta Fairmaire, 1899 Synapsis Bates, 1868 Synchilus Britton, 1956 Synclitopa Kolbe, 1897 Synenaria Lacroix, 1993 Syngeneschiza Brenske, 1898 Synistovalgus Kolbe, 1897 Syrichthodontus Péringuey, 1901 Syrichthomorphus Péringuey, 1901 Syrichthoschema Janssens, 1942 Systellopus Sharp, 1877 Systellorrhina Kraatz, 1895 Taeniodera Burmeister, 1842 Tafaia Valck Lucassens, 1939 Taiwanotrichia Kobayashi, 1990 Talautoclyster Yamaya, 2001 Tamnoserica Brenske, 1899 Tanyana Stebnicka, 2006 Tanyproctoides Petrovitz, 1971 Tanyproctus Ménétriés, 1832 Tanzanilepis Lacroix, 2008 Tanzanipholis Lacroix, 2002 Tanzanolus Scholtz & Howden, 1987 Taphrocephala Quedenfeldt, 1888 Tapinoschema Thomson, 1880 Tarsocamenta Moser, 1924 Tarsovalgus Miyake, 1993 Taurhina Burmeister, 1842 Tehuacania Endrödi, 1975 Teinogenys Sharp, 1873 Telaugis Burmeister, 1844 Telochilus Krikken, 1975 Telura Erichson, 1842 Teluroides Britton, 1990 Temnoplectron Westwood, 1841 Temnorhynchus Hope, 1837 Tephraea Burmeister, 1842 Tephraeoserica Brenske, 1900 Teraserica Brenske, 1897 Terebrogaster Lacroix, 1989 Termitaxis Krikken, 1970 Termitoderus Mateu, 1966 Termitodiellus Nakane, 1961 Termitodius Wasmann, 1894 Termitophilus Britton, 1957 Termitopisthes Wasmann, 1918 Termitotrox Reichensperger, 1915 Territonia Krikken, 2018 Tesarius Rakovič, 1981 Tesserodon Hope, 1837 Tesserodoniella Vaz-de-Mello & Halffter, 2006 Testaceitrichia Matsumoto, 2021 Tetraclipeoides Schmidt, 1914 Tetraechma Blanchard, 1845 Tetragonorrhina Kraatz, 1896 Tetragus Duarte & Grossi, 2022 Tetraodorhina Blanchard, 1842 Tetraserica Ahrens, 2004 Teuchestes Mulsant, 1842 Thabina Péringuey, 1902 Theodosia Thomson, 1880 Theotimius Huchet, 2000 Theuremaripa Soula, 2006 Thinorycter Semenov & Reichardt, 1925 Thoracoplia Prokofiev, 2015 Thoracotrichia Decelle, 1968 Thronistes Burmeister, 1847 Thrymoserica Brenske, 1897 Thyce LeConte, 1856 Thyregis Blackburn, 1904 Thyreogonia Reitter, 1898 Thyridium Burmeister, 1844 Thyriochlorota Ohaus, 1915 Tiamidema Prokofiev, 2014 Tiarocera Burmeister, 1842 Tiaronthophagus Roggero, Moretto, Barbero & Palestrini, 2019 Tibiodrepanus Krikken, 2009 Tibiotrichius Miyake, 1994 Tibiovalgus Krikken, 1978 Tiniocellus Péringuey, 1901 Tipicha Soula, 2002 Tlaocera Péringuey, 1904 Tmesorrhina Westwood, 1841 Tocama Reitter, 1902 Tomarus Erichson, 1847 Tomogonus Orbigny, 1904 Tonkinilla Prokofiev, 2021 Torynorrhina Arrow, 1907 Tosevskiana Pavicevic, 1985 Toxocnemis Dombrow, 2002 Toxospathius Fairmaire, 1878 Trachyserica Brenske, 1899 Tragiscus Klug, 1855 Trematodes Faldermann, 1835 Triacmoserica Kolbe, 1914 Tribialus Dupuis & Perrin, 2022 Tribopertha Reitter, 1903 Trichanomala Arrow, 1917 Trichaphodiellus Schmidt, 1914 Trichaphodioides Paulian, 1942 Trichaphodius Schmidt, 1914 Trichaulax Kraatz, 1880 Trichesthes Erichson, 1847 Trichillidium Vaz-de-Mello, 2008 Trichillum Harold, 1868 Trichinopus Waterhouse, 1875 Trichioaparammoecius Minkina, 2018 Trichiodera Burmeister, 1855 Trichiopsammobius Petrovitz, 1963 Trichiorhyssemus Clouët, 1901 Trichiotinus Casey, 1915 Trichius Fabricius, 1775 Trichocephala Moser, 1916 Trichogomphus Burmeister, 1847 Tricholepis Blanchard, 1851 Tricholontha Nomura, 1952 Trichomaladera Nomura, 1974 Trichoneptunides Legrand, 2001 Trichonotuloides Balthasar, 1945 Trichonotulus Bedel, 1911 Trichoparadidactylia Bordat, 2022 Trichoplus Burmeister, 1842 Trichopopillia Ohaus, 1897 Trichoschiza Moser, 1917 Trichostetha Burmeister, 1842 Trigonochilus Brenske, 1896 Trigonocnemis Kraatz, 1894 Trigonopeltastes Burmeister & Schaum, 1840 Trigonophorinus Pouillaude, 1913 Trigonophorus Hope, 1831 Trigonoscelus (Petrovitz, 1963) Trigonostomus Brenske, 1893 Trinoxia Brenske, 1894 Triodontella Reitter, 1919 Triodontonyx Moser, 1921 Triodontus Westwood, 1845 Triodonyx Saylor, 1942 Trioplognathus Ohaus, 1904 Trioplus Burmeister, 1847 Trioserica Moser, 1922 Triplognatha Krikken, 1987 Trissodon Burmeister, 1847 Tristaphodius Balthasar, 1932 Trizogeniates Ohaus, 1917 Trochaloschema Reitter, 1896 Trochaloserica Brenske, 1900 Trochalus Laporte, 1832 Trogodes Boheman, 1857 Tropidonitis Janssens, 1937 Tropinota Mulsant, 1842 Tropiorhynchus Blanchard, 1851 Trymodera Gerstaecker, 1867 Trypoxylus Minck, 1920 Tsaratanoplia Lacroix, 1997 Tulbaghia Péringuey, 1904 Turanella Semenov, 1905 Uenoeus Ochi & Kon, 2017 Ulata Saylor, 1945 Uloptera Burmeister, 1842 Unidentis Josso & Prévost, 2008 Upsa Deschodt, Sole & Scholtz, 2020 Uroxys Westwood, 1842 Vadonaria De Wailly, 1950 Vadonidella Ruter, 1973 Valgoides Fairmaire, 1899 Valgus Scriba, 1790 Vansoniola Schein, 1958 Varencya Lacroix, 1993 Vayana Ohaus, 1915 Versicorpus Deschodt, Davis & Scholtz, 2011 Vezoanus Lacroix, 1993 Vieuella Ruter, 1964 Viridimicus Jameson, 1990 Vladimirellus Dellacasa, Dellacasa & Bordat, 2002 Volinus Mulsant & Rey, 1869 Wadaia Itoh, 1993 Walsternoplus Allsopp, Jákl & Rey, 2023 Walterantus Cambefort, 1977 Wambo Allsopp, 1988 Warwickia Smith & Evans, 2005 Watkinsia Britton, 1995 Webbella Britton, 1988 Wernoryctes Dechambre, 1999 Xanthotrogus Reitter, 1902 Xeloma Kraatz, 1881 Xenaclopus Arrow, 1915 Xenocanthon Martinez, 1952 Xenoceraspis Arrow, 1920 Xenochlorota Soula, 2002 Xenodorus Brême, 1844 Xenogeniates Villatoro & Jameson, 2001 Xenoheptaulacus Hinton, 1934 Xenoloba Bates, 1889 Xenopelidnota Bates, 1904 Xenoproctis Kolbe, 1896 Xenoreoderus Arrow, 1910 Xenoserica Ahrens, 2005 Xenotrochalus Moser, 1917 Xenotrogus Britton, 1978 Xeropsamobeus Saylor, 1937 Xinidium Harold, 1869 Xiphoscelis Burmeister, 1842 Xiphosceloides Holm, 1992 Xylonichus Boisduval, 1835 Xyloryctes Hope, 1837 Xyloscaptes Prell, 1934 Xylotrupes Hope, 1837 Xyridea Britton, 1990 Xyrine Britton, 1987 Xyroa Britton, 1987 Xyrodes Britton, 1987 Yaaxkumukia Morón & Nogueira, 1998 Yanovalgus Nomura, 1952 † Yixianscarabaeus Nikolajev, 2015 Youngaphodius Bordat, 1988 Yvescambefortius Ochi & Kon, 1996 Zaburina Saylor, 1945 Zagurya Fuhrmann, 2021 Zebinus Fairmaire, 1894 Zietzia Blackburn, 1894 Zimbabuephylla Lacroix, 1997 Zomba Lacroix, 2006 Zonocopris Arrow, 1932 Les scarabées dans la culture
Collection d'amulettes en forme de scarabées. Le Scarabée sacré (Scarabaeus sacer ) est un animal vénéré dans l'Égypte antique. Symbole du soleil et de Rê, il était l'un des douze animaux sacrés associé aux douze heures du jour et de la nuit[réf. nécessaire] .
Le Scarabée rhinocéros siamois ou scarabée de combat (Xylotrupes gideon) est capturé dans la nature puis dressé pour des duels qui font l'objet de paris dans le nord de la Thaïlande. Ces duels de scarabées sont couramment organisés. Ces combats d’insectes du type xylotrupes appelés Kwang localement sont à bien des égards surprenants, notamment du fait qu’il est sans doute plus difficile d’attribuer à un scarabée qu’à un autre animal des caractéristiques anthropomorphiques et qu'il est plus compliqué encore de deviner ce qu’il perçoit. Le duel de scarabées met à l’épreuve une forme de zoomanité trouble fondée sur une forme d’attachement inédite entre l’homme et l’animal, .
Notes et références