cetaceans
Cetacea (/sɪˈteɪʃə/; from Latin cetus 'whale', from Ancient Greek κῆτος (kêtos) 'huge fish, sea monster') is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins, and porpoises.
The Cetaceans Portal
Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver.
While the majority of cetaceans live in marine environments, a small number reside solely in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons.
Cetaceans are famous for their high intelligence, complex social behaviour, and the enormous size of some of the group's members. For example, the blue whale reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short tons), making it the largest animal known ever to have existed.
There are approximately 89 living species split into two parvorders: Odontoceti or toothed whales (containing porpoises, dolphins, other predatory whales like the beluga and the sperm whale, and the poorly understood beaked whales) and the filter feeding Mysticeti or baleen whales (which includes species like the blue whale, the humpback whale and the bowhead whale). Despite their highly modified bodies and carnivorous lifestyle, genetic and fossil evidence places cetaceans as nested within even-toed ungulates, most closely related to hippopotamus within the clade Whippomorpha. (Full article...)
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A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors.
In 1859, when Charles Darwin's On the Origin of Species was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be urged against my theory," but he explained it by relating it to the extreme imperfection of the geological record. He noted the limited collections available at the time but described the available information as showing patterns that followed from his theory of descent with modification through natural selection. Indeed, Archaeopteryx was discovered just two years later, in 1861, and represents a classic transitional form between earlier, non-avian dinosaurs and birds. Many more transitional fossils have been discovered since then, and there is now abundant evidence of how all classes of vertebrates are related, including many transitional fossils. Specific examples of class-level transitions are: tetrapods and fish, birds and dinosaurs, and mammals and "mammal-like reptiles". (Full article...) -
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. Herman Melville often referred to whales as "Leviathans" in his book. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales (or "raptorial sperm whales") and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Livyatan's total length has been estimated to be about 13.5–17.5 m (44–57 ft), almost similar to that of the modern sperm whale (Physeter macrocephalus), making it one of the largest predators known to have existed. The teeth of Livyatan measured 36.2 cm (1.19 ft), and are the largest biting teeth of any known animal, excluding tusks. It is distinguished from the other raptorial sperm whales by the basin on the skull spanning the length of the snout. The spermaceti organ contained in that basin is thought to have been used in echolocation and communication, or for ramming prey and other sperm whales. The whale may have interacted with the large extinct shark megalodon (Otodus megalodon), competing with it for a similar food source. Its extinction was probably caused by a cooling event at the end of the Miocene period causing a reduction in food populations. The geological formation where the whale has been found has also preserved a large assemblage of marine life, such as sharks and marine mammals. (Full article...) -
Species of the infraorder Cetacea
The evolution of cetaceans is thought to have begun in the Indian subcontinent from even-toed ungulates (Artiodactyla) 50 million years ago (mya) and to have proceeded over a period of at least 15 million years. Cetaceans are fully aquatic marine mammals belonging to the order Artiodactyla and branched off from other artiodactyls around 50 mya. Cetaceans are thought to have evolved during the Eocene (56-34 mya), the second epoch of the present-extending Cenozoic Era. Molecular and morphological analyses suggest Cetacea share a relatively recent closest common ancestor with hippopotami and that they are sister groups. Being mammals, they surface to breathe air; they have 5 finger bones (even-toed) in their fins; they nurse their young; and, despite their fully aquatic life style, they retain many skeletal features from their terrestrial ancestors. Research conducted in the late 1970s in Pakistan revealed several stages in the transition of cetaceans from land to sea.
The two modern parvorders of cetaceans – Mysticeti (baleen whales) and Odontoceti (toothed whales) – are thought to have separated from each other around 28–33 mya in a second cetacean radiation, the first occurring with the archaeocetes. The adaptation of animal echolocation in toothed whales distinguishes them from fully aquatic archaeocetes and early baleen whales. The presence of baleen in baleen whales occurred gradually, with earlier varieties having very little baleen, and their size is linked to baleen dependence (and subsequent increase in filter feeding). (Full article...) -
The fin whale (Balaenoptera physalus), also known as the finback whale or common rorqual, is a species of baleen whale and the second-longest cetacean after the blue whale. The biggest individual reportedly measured 26 m (85 ft) in length, with a maximum recorded weight of 77,000–81,000 kg (170,000–179,000 lb). The fin whale's body is long, slender and brownish-gray in color, with a paler underside to appear less conspicuous from below (countershading).
At least two recognized subspecies exist, one in the North Atlantic and one across the Southern Hemisphere. It is found in all the major oceans, from polar to tropical waters, though it is absent only from waters close to the pack ice at the poles and relatively small areas of water away from the open ocean. The highest population density occurs in temperate and cool waters. Its prey mainly consists of smaller schooling fish, small squid, or crustaceans, including copepods and krill. Mating takes place in temperate, low-latitude seas during the winter. Fin whales are often observed in pods of 6–10 animals, with whom they communicate utilizing frequency-modulated sounds, ranging from 16 to 40 hertz. (Full article...) -
Pilot whales are cetaceans belonging to the genus Globicephala. The two extant species are the long-finned pilot whale (G. melas) and the short-finned pilot whale (G. macrorhynchus). The two are not readily distinguishable at sea, and analysis of the skulls is the best way to distinguish between the species. Between the two species, they range nearly worldwide, with long-finned pilot whales living in colder waters and short-finned pilot whales living in tropical and subtropical waters. Pilot whales are among the largest of the oceanic dolphins, exceeded in size only by the orca. They and other large members of the dolphin family are also known as blackfish.
Pilot whales feed primarily on squid, but will also hunt large demersal fish such as cod and turbot. They are highly social and may remain with their birth pod throughout their lifetime. Short-finned pilot whales are one of the few mammal species in which females go through menopause, and postreproductive females continue to contribute to their pod. Pilot whales are notorious for stranding themselves on beaches, but the reason behind this is not fully understood, although marine biologists have shed light on the discovery it is due to the mammals inner ear (their principal navigational sonar) being damaged from noise-pollution in the ocean, such as from cargo ships or military exercises. The conservation status of short-finned and long-finned pilot whales has been determined to be least concern. (Full article...) -
The sperm whale or cachalot (Physeter macrocephalus) is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus Physeter and one of three extant species in the sperm whale family, along with the pygmy sperm whale and dwarf sperm whale of the genus Kogia.
The sperm whale is a pelagic mammal with a worldwide range, and will migrate seasonally for feeding and breeding. Females and young males live together in groups, while mature males (bulls) live solitary lives outside of the mating season. The females cooperate to protect and nurse their young. Females give birth every four to twenty years, and care for the calves for more than a decade. A mature sperm whale has no natural predators, although calves and weakened adults are sometimes killed by pods of killer whales (orcas). (Full article...) -
Whale barnacles are species of acorn barnacle that belong to the family Coronulidae. They typically attach to baleen whales, and sometimes settle on toothed whales. The whale barnacles diverged from the turtle barnacles about three million years ago.
Whale barnacles passively filter food, using tentacle-like cirri, as the host swims through the water. The arrangement is generally considered commensal as it is done at no cost or benefit to the host. However, some whales may make use of the barnacles as protective armor or for inflicting more damage while fighting, which would make the relationship mutualistic where both parties benefit; alternatively, some species may just increase the drag that the host experiences while swimming, making the barnacles parasites. (Full article...) - Cetacea is an infraorder that comprises the 94 species of whales, dolphins, and porpoises. It is divided into toothed whales (Odontoceti) and baleen whales (Mysticeti), which diverged from each other in the Eocene some 50 million years ago (mya). Cetaceans are descended from land-dwelling hoofed mammals, and the now extinct archaeocetes represent the several transitional phases from terrestrial to completely aquatic. Historically, cetaceans were thought to have descended from the wolf-like mesonychians, but cladistic analyses confirm their placement with even-toed ungulates in the order Cetartiodactyla.
Whale populations were drastically reduced in the 20th century from intensive whaling, and the activity was globally banned in 1982. Smaller cetaceans are at risk of accidentally getting caught by fishing vessels using, namely, seine fishing, drift netting, or gill netting operations. (Full article...) -
Janjucetus is an extinct genus of cetacean, and a basal baleen whale (Mysticeti), from the Late Oligocene around 25 million years ago (mya) off south-east Australia, containing one species J. hunderi. Unlike modern mysticetes, it possessed large teeth for gripping and shredding prey, and lacked baleen, and so was likely to have been a predator that captured large single prey animals rather than filter feeding. However, its teeth may have interlocked, much like those of the modern-day filter-feeding crabeater seal (Lobodon carcinophaga), which would have allowed some filter-feeding behaviour. Its hunting behaviour was probably similar to the modern-day leopard seal (Hydrurga leptonyx), probably eating large fish. Like baleen whales, Janjucetus could not echolocate; however, it did have unusually large eyes, and so probably had an acute sense of vision. The only specimen was found on the Jan Juc beach, where the remains of the extinct whales Mammalodon, Prosqualodon and Waipatia have also been discovered. (Full article...) -
Arktocara is an extinct genus of river dolphin from the Oligocene epoch of Alaska, containing one species, A. yakataga. Having been discovered in 25-million-year-old strata near the 60th parallel north, it is perhaps the oldest-known crown toothed whale and the northmost river dolphin discovered. It was a member of the now-extinct family Allodelphinidae, along with the genera Allodelphis, Goedertius, Ninjadelphis, and Zarhinocetus. It measured approximately 2.26 or 2.28 meters (7.4 or 7.5 ft), comparable to its closest living relative, the South Asian river dolphin, which measures 2.4 meters (7.9 ft). However, the animal probably had an elongated beak and neck, so it may have been longer. The animal is known only from a partially preserved skull. Its ecology may have been similar to the modern-day Dall's porpoise, and it may have competed with contemporaneous delphinoids. Its remains were found in the Poul Creek Formation, which has also yielded several mollusk species. (Full article...) -
The anatomist John Struthers (at left, in top hat) with the Tay Whale at John Woods' yard, Dundee, 1884, photographed by George Washington Wilson
The Tay Whale, known locally as the Monster, was a humpback whale that swam into the Firth of Tay of eastern Scotland in 1883. It was harpooned in a hunt, but escaped, and was found floating dead off Stonehaven a week later. It was towed into Dundee by a showman, John Woods, and exhibited on a train tour of Scotland and England.
The Regius Professor of Anatomy at Aberdeen University, John Struthers dissected the whale, much of the time in public with a military band playing in the background, organised by Woods. The decomposing whale made Woods a great deal of money, and Struthers famous. (Full article...) -
Orcinus citoniensis is an extinct species of killer whale identified in the Late Pliocene of Italy and the Early Pleistocene of England. It was smaller than the modern killer whale (O. orca), 4 m (13 ft) versus 7 to 10 m (23 to 33 ft), and had around 8 more teeth in its jaw. It may have resembled the modern killer whale in appearance, and could represent a transitional species between the modern killer whale and other dolphins. O. citoniensis could have hunted fish and squid in pods, and coexisted with other large predators of the time such as the orcinine Hemisyntrachelus and the extinct shark Otodus megalodon. (Full article...) -
Brygmophyseter, known as the biting sperm whale, is an extinct genus of toothed whale in the sperm whale family with one species, B. shigensis. When it was first described in 1994, the species was placed in the genus Scaldicetus based on tooth morphology, but this was later revised in 1995. In 2006, it was classified into the genus Naganocetus, which is considered to be a junior synonym. The only known specimen, a nearly complete skeleton, was dated to be around 16–15 million years old (middle Miocene). Brygmophyseter is thought to have been 6.5–7 meters (21–23 ft) long, and it probably had 11 or 12 teeth in the upper and lower jaws. Brygmophyseter is part of a group of macroraptorial sperm whales (often shortened to "raptorial") which tended to be apex predators using their large teeth to catch struggling prey such as whales. It had a spermaceti organ which was probably used for biosonar like in the modern sperm whale. The whale has made an appearance on The History Channel's TV series Jurassic Fight Club. (Full article...) -
Ambulocetus (Latin ambulare "to walk" + cetus "whale") is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, Ambulocetus natans (Latin natans "swimming"), known solely from a near-complete skeleton. Ambulocetus is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. Ambulocetus is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes Himalayacetus and Gandakasia (also from the Eocene of the Indian subcontinent).
Ambulocetus had a narrow, streamlined body, and a long, broad snout, with eyes positioned at the very top of its head. Because of these features, it is hypothesised to have behaved much like a crocodile, waiting near the water's surface to ambush large mammals, using its powerful jaws to clamp onto and drown or thrash prey. Additionally, its ears possessed similar traits to modern cetaceans, which are specialised for hearing and detecting certain frequencies underwater, although it is unclear if Ambulocetus also used these specialised ears for hearing underwater. They may have instead been utilised for bone conduction on land, or perhaps served no function for early cetaceans. (Full article...) -
Zygophyseter varolai is an extinct sperm whale that lived during the Tortonian age of the Late Miocene 11.2 to 7.6 million years ago. It is known from a single specimen from the Pietra Leccese Formation in Italy. It was a member of a stem group of fossil macroraptorial sperm whales (often shortened to "raptorial") also including Brygmophyseter, Acrophyseter, and Livyatan. It probably grew to be around 6.5 to 7 meters (21 to 23 ft) in length and shared some characteristics with other raptorials, such as large teeth with tooth enamel that were functional in both the upper and lower jaws which the modern sperm whale (Physeter macrocephalus) lacks. It also had a beak, the ability to echolocate prey, and could have probably swum faster than the modern-day sperm whale which can reach 4 kilometers per hour (2.5 mph). These were probably used in the capture of large prey, such as large fish, seals, and whales. In fact, its common name, the killer sperm whale, refers to its feeding habits that would have had a resemblance to the modern-day killer whale (Orcinus orca). (Full article...)
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Photo credit: Captain Budd Christman of the NOAA.
The Humpback Whale (Megaptera novaeangliae) is a mammal which belongs to the baleen whale suborder. It is a large whale: an adult usually ranges between 12–16 m (40–50 ft) long and weighs approximately 36,000 kilograms (79,000 lb). It is well known for its breaching (leaping out of the water), its unusually long front fins, and its complex whale song. The Humpback Whale lives in oceans and seas around the world, and is regularly sought out by whale-watchers.
More did you know...
- ...common dolphins, which are often seen off South Africa’s east coast, can occur in schools of several thousand. The biggest school on record was estimated to consist of about 15,000 dolphins!
- ...because whales and dolphins are streamlined to swim in water, they do not have external organs. This makes it almost impossible to tell the sex of a whale or dolphin when watching them on the sea surface.
- ...there are probably types of cetaceans that are as yet unknown. For example, the Longman's beaked whale is only known from skulls washed ashore in Somalia and Australia. It has never been seen alive!
- ...in baleen whales females are bigger than males, while in the toothed whales the males are bigger than females.
- ...the open mouth of an adult southern right whale can be two metres wide!
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General images -
The following are images from various cetacean-related articles on Wiki English.
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Bryde's whale (rorqual) (from Baleen whale) -
Their eyes are relatively small for their size. -
Fossil of Squalodon (from Toothed whale) -
Eden's whale illustration -
A protest against Japan's scientific whaling -
Female right whale with calf -
Archaeomysticetes, like Janjucetus, had teeth. -
Humpback whale illustration -
Gray whale illustration -
Diagram illustrating sound generation, propagation and reception in a toothed whale. Outgoing sounds are red and incoming ones are green (from Toothed whale) -
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Atlantic white-sided dolphin caught in a drive hunt in Hvalba on the Faroe Islands being taken away with a forklift (from Toothed whale) -
Japanese scientific whaling on a pair of Antarctic minke whales (from Baleen whale) -
Archaeocetes (like this Basilosaurus) had a heterodont dentition (from Evolution of cetaceans) -
Orca hunting a Weddell seal (from Toothed whale) -
Baleen whales vary considerably in size and shape, depending on their feeding behavior. (from Baleen whale) -
Acrophyseter skull (from Evolution of cetaceans) -
South Atlantic right whale illustration -
Accessory baleen plates taper off into small hairs (from Baleen whale) -
A humpback whale skeleton. Notice how the jaw is split into two. -
Species of the infraorder Cetacea (from Evolution of cetaceans) -
North Atlantic right whale illustration -
Biosonar by cetaceans (from Toothed whale) -
Bowhead whale illustration -
Dolphins (aquatic mammals) and ichthyosaurs (extinct marine reptiles) share a number of unique adaptations for fully aquatic lifestyle and are frequently used as extreme examples of convergent evolution (from Evolution of cetaceans) -
Features of a sperm whale skeleton (from Toothed whale) -
The skeleton of a bowhead whale with the hind limb and pelvic bone structure circled in red. This bone structure stays internal during the entire life of the species. (from Evolution of cetaceans) -
Spectrogram of dolphin vocalizations. Whistles, whines, and clicks are visible as upside down V's, horizontal striations, and vertical lines, respectively. (from Toothed whale) -
Anatomy of the bottlenose dolphin (from Toothed whale) -
Fin whale illustration -
The heart of a blue whale with a person standing next to it -
Common minke whale illustration -
Orange whale lice on a right whale -
The nose of the whale is filled with a waxy substance that was widely used in candles, oil lamps, and lubricants (from Toothed whale) -
Pygmy right whale illustration -
Possible relationships between cetaceans and other ungulate groups. (from Evolution of cetaceans) -
Blue whale illustration -
The remains of a North Atlantic right whale after it collided with a ship propeller -
Antarctic minke whale illustration -
A phylogenetic tree showing the relationships among cetacean families. (from Evolution of cetaceans) -
Pakicetus attocki skeleton (from Evolution of cetaceans) -
Sei whale illustration -
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An orca by the name of Ulises performing at SeaWorld, 2009 (from Toothed whale) -
Humpback whales lunge-feeding in the course of bubble net fishing -
Megalodon hunted Miocene baleen whales (from Baleen whale) -
World population graph of blue whales -
Restoration of Janjucetus hunderi (from Baleen whale)
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Did you know (auto-generated)
- ... that one of the first researchers to propose dolphin-assisted therapy for humans later renounced it?
- ... that the South Asian river dolphin is nearly blind and relies on echolocation for navigation?
- ... that Celia Kaye won the Golden Globe Award for Most Promising Newcomer in 1965 for her starring role in Island of the Blue Dolphins?
- ... that one can swim with humpback whales in the Niue Nukutuluea Multiple-Use Marine Park?
- ... that the three dolphins on the coat of arms of Anguilla represent "unity, strength and endurance", which is also the motto of the territory?
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Cetacean articles
| Whale species | Andrews' Beaked Whale • Balaenoptera omurai • Beluga • Blainville's Beaked Whale • Blue Whale |
| Dolphin species | Atlantic Spotted Dolphin • Atlantic White-sided Dolphin • Australian Snubfin Dolphin • Baiji • Boto • Chilean Dolphin • Clymene Dolphin • Commerson's Dolphin • Common Bottlenose Dolphin • Dusky Dolphin |
| Porpoise species | Burmeister's Porpoise • Dall's Porpoise • Finless Porpoise • Harbour Porpoise • Spectacled Porpoise • Vaquita |
| Other articles | Aboriginal whaling • Ambergris • Animal echolocation • Archaeoceti • Baleen • Baleen whale • Beached whale • Beaked Whale • Blowhole (biology) • Blubber • Bottlenose dolphin |
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