Bird Bunting

The buntings are a group of Old World passerine birds forming the genus Emberiza, the only genus in the family Emberizidae. The family contains 45 species. They are seed-eating birds with stubby, conical bills.

Bunting
Bird Bunting
Cirl bunting (Emberiza cirlus)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Superfamily: Emberizoidea
Family: Emberizidae
Vigors, 1831
Genus: Emberiza
Linnaeus, 1758
Type species
Emberiza citrinella
Species

44, see text

Synonyms
  • Onychospina Bonaparte, 1853
  • Onychospiza Rey, 1872 (incorrect subsequent spelling)

Taxonomy

The family Emberizidae was formerly much larger and included the species now placed in the Passerellidae (New World sparrows) and Calcariidae (longspurs and snow buntings). Molecular phylogenetic studies found that the large family consisted of distinct clades that were better treated as separate families.

The genus Emberiza is now the only genus placed in the family Emberizidae. The genus was introduced by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae. The type species was subsequently designated as the yellowhammer (Emberiza citrinella). The genus name Emberiza is from Old German Embritz, a bunting. The origin of the English "bunting" is unknown.

A 2008 genetic study found that three emberizid species that were placed in their own monotypic genera clustered within the Emberiza. These were the crested bunting (Melophus lathami), the slaty bunting (Latouchiornis siemsseni), and the corn bunting (Miliaria calandra). All three species are now included in the genus Emberiza.

A large DNA-based study of the passerines published in 2019 found that the buntings are most closely related to the longspurs and snow buntings in the family Calcariidae.

Ornithologists Edward Dickinson and Leslie Christidis in the fourth edition of the Howard and Moore Complete Checklist of the Birds of the World chose to split up Emberiza and recognise the genera Fringillaria, Melophus, Granativora, Emberiza, and Schoeniclus. Their example has not been followed by the online version of the Handbook of the Birds of the World nor by Frank Gill and David Donsker in the list of world birds that they maintain on behalf of the International Ornithologists' Union. The British Ornithologists' Union has argued that splitting the genus provides little benefit and destabilizes the nomenclature.

Species in the New World genus Passerina include the word "bunting" in their common names, but are now classed in the family Cardinalidae.

The family is divided into four major clades. The species in Clade I are mainly African while those in Clades II to IV are Palearctic:

Emberizidae
Clade I

Cabanis's bunting – Emberiza cabanisi

Golden-breasted bunting – Emberiza flaviventris

Somali bunting – Emberiza poliopleura

Cape bunting – Emberiza capensis

Lark-like bunting – Emberiza impetuani

Socotra bunting – Emberiza socotrana

Gosling's bunting – Emberiza goslingi

Cinnamon-breasted bunting – Emberiza tahapisi

House bunting – Emberiza sahari

Striolated bunting – Emberiza striolata

Clade II

Yellow-throated bunting – Emberiza elegans

Slaty bunting – Emberiza siemsseni

Ochre-rumped bunting – Emberiza yessoensis

Pallas's bunting – Emberiza pallasi

Common reed bunting – Emberiza schoeniclus

Yellow-browed bunting – Emberiza chrysophrys

Tristram's bunting – Emberiza tristrami

Grey bunting – Emberiza variabilis

Yellow-breasted bunting – Emberiza aureola

Chestnut bunting – Emberiza rutila

Little bunting – Emberiza pusilla

Rustic bunting – Emberiza rustica

Black-faced bunting – Emberiza spodocephala

Yellow bunting – Emberiza sulphurata

Clade III

Crested bunting – Emberiza lathami

Red-headed bunting – Emberiza bruniceps

Black-headed bunting – Emberiza melanocephala

Clade IV

Corn bunting – Emberiza calandra

Chestnut-eared bunting – Emberiza fucata

Tibetan bunting – Emberiza koslowi

Jankowski's bunting – Emberiza jankowskii

Meadow bunting – Emberiza cioides

Rock bunting – Emberiza cia

Godlewski's bunting – Emberiza godlewskii

Grey-necked bunting – Emberiza buchanani

Cinereous bunting – Emberiza cineracea

Cretzschmar's bunting – Emberiza caesia

Ortolan bunting – Emberiza hortulana

Cirl bunting – Emberiza cirlus

White-capped bunting – Emberiza stewarti

Yellowhammer – Emberiza citrinella

Pine bunting – Emberiza leucocephalos

The above cladogram is based on a study published in 2021. The phylogenetic relationships of two African species, the brown-rumped bunting (Emberiza affinis) and Vincent's bunting (Emberiza vincenti), were not determined.

List of species

The genus contains 44 species.

Image Common Name Scientific name Distribution
Bird Bunting  Crested bunting Emberiza lathami Southeast Asia.
Slaty bunting Emberiza siemsseni China.
Bird Bunting  Corn bunting Emberiza calandra Western Europe and North Africa across to northwestern China.
Bird Bunting  Yellowhammer Emberiza citrinella southeast England and most of Europe east to the northwestern corner of Russia and western Ukraine.
Bird Bunting  Pine bunting Emberiza leucocephalos Asia
Bird Bunting  Rock bunting Emberiza cia northwest Africa, southern Europe east to central Asia, and the Himalayas
Bird Bunting  Godlewski's bunting Emberiza godlewskii China, Pakistan, India, Kazakhstan, Mongolia, Myanmar, and Russia.
Bird Bunting  Meadow bunting Emberiza cioides southern Siberia, northern and eastern China, eastern Kazakhstan, Kyrgyzstan, Mongolia, Korea and Japan.
Bird Bunting  White-capped bunting Emberiza stewarti Afghanistan, India, Iran, Kazakhstan, Kyrgyzstan, Nepal, Pakistan, Tajikistan, Turkmenistan, and Uzbekistan.
Jankowski's bunting Emberiza jankowskii Russian Far East, Manchuria and far northeastern Korea
Bird Bunting  Grey-necked bunting Emberiza buchanani Caspian Sea to the Altai Mountains in Central Asia
Bird Bunting  Cinereous bunting Emberiza cineracea southern Turkey and southern Iran
Bird Bunting  Ortolan bunting Emberiza hortulana European countries and western Asia
Bird Bunting  Cretzschmar's bunting Emberiza caesia Greece, Turkey, Cyprus and the Levant.
Bird Bunting  Cirl bunting Emberiza cirlus southern Europe, on the Mediterranean islands and in north Africa
Bird Bunting  Striolated bunting Emberiza striolata Chad, east through south-west Asia to north-western India, Africa
Bird Bunting  House bunting Emberiza sahari northwestern Africa from Morocco south to Mali and east to Chad.
Bird Bunting  Lark-like bunting Emberiza impetuani Angola, Botswana, Lesotho, Namibia, South Africa, and Zimbabwe.
Bird Bunting  Cinnamon-breasted bunting Emberiza tahapisi mainland sub-Saharan Africa
Bird Bunting  Gosling's bunting Emberiza goslingi Mauritania and Senegal to south-western Sudan and north-eastern Democratic Republic of Congo.
Socotra bunting Emberiza socotrana Yemen
Bird Bunting  Cape bunting Emberiza capensis southern Africa from south-western Angola, eastern Zambia, Zimbabwe and southern Tanzania to the Cape.
Bird Bunting  Tristram's bunting Emberiza tristrami eastern Manchuria and the Russian Far East and winters in central and southern China.
Bird Bunting  Chestnut-eared bunting Emberiza fucata the Himalayas locally across China to south-eastern Siberia, Korea and northern Japan
Bird Bunting  Little bunting Emberiza pusilla north-east of Europe and northern Eurosiberia to the Russian Far East and northern India, southern China and the northern parts of south-east Asia.
Bird Bunting  Yellow-browed bunting Emberiza chrysophrys eastern Siberia, China
Bird Bunting  Rustic bunting Emberiza rustica south-east Asia, Japan, Korea, and eastern China.
Bird Bunting  Yellow-throated bunting Emberiza elegans China, Japan, Korea, Myanmar, Russia, and Taiwan.
Bird Bunting  Yellow-breasted bunting Emberiza aureola Finland to Bering Sea migrating to Indochina
Bird Bunting  Somali bunting Emberiza poliopleura Ethiopia, Kenya, Somalia, South Sudan, Tanzania, and Uganda
Bird Bunting  Golden-breasted bunting Emberiza flaviventris Africa south of the Sahara
Bird Bunting  Brown-rumped bunting Emberiza affinis Senegal to Sudan and Uganda
Bird Bunting  Cabanis's bunting Emberiza cabanisi sub-Saharan Africa
Bird Bunting  Chestnut bunting Emberiza rutila Siberia, northern Mongolia and north-eastern China.
Tibetan bunting Emberiza koslowi Tibet
Bird Bunting  Black-headed bunting Emberiza melanocephala Japan, China, Hong Kong, Thailand, Laos, South Korea and Malaysia
Bird Bunting  Red-headed bunting Emberiza bruniceps Asia-Afghanistan, Iran, Kazakhstan, Kyrgyzstan, Mongolia; Russian Federation (European Russia, Central Asian Russia), Tajikistan, Turkmenistan, Uzbekistan.
Bird Bunting  Yellow bunting Emberiza sulphurata Japan
Bird Bunting  Black-faced bunting Emberiza spodocephala southern Siberia across to northern China.
Bird Bunting  Masked bunting Emberiza personata Sakhalin, the Kuril Islands, and Japan.
Bird Bunting  Grey bunting Emberiza variabilis Kamchatka, Sakhalin, Kuril Islands and northern Japan
Bird Bunting  Pallas's reed bunting Emberiza pallasi northeast European Russia, north Kamchatka
Bird Bunting  Ochre-rumped bunting Emberiza yessoensis eastern Mongolia, northeast China and Ussuriland
Bird Bunting  Common reed bunting Emberiza schoeniclus Europe

Extinct species have been described:

  • Long-legged bunting (Emberiza alcoveri) (Late Quaternary)
  • Emberiza shaamarica (Late Pliocene of Central Asia)
  • Emberiza polgardiensis (Upper Miocene of Hungary)
  • Emberiza media (Pliocene of Hungary)
  • Emberiza parva (Pliocene of Hungary)
  • Emberiza gaspariskii (Pliocene of Hungary)
  • Emberiza bartoki (Middle Miocene of Hungary) (Nomen dubium)

Emberiza pannonica from the upper Miocene of Hungary is also referred to this genus, but was later found to be a member of Muscicapidae.

References

Further reading

Buntings and Sparrows - A Guide to the Buntings and North American Sparrows by Urban Olsson and Jon Curson, illustrated by Clive Byers (1995) ISBN 1-873403-19-4

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Bird Bunting TaxonomyBird Bunting List of speciesBird Bunting Further readingBird Bunting

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